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Morning glory (Ipomoea tricolor) flower with pollen grains scattered by visiting insects
Common rue (Ruta graveolens) flower in visible light, nectar at base of petals. Stamens flip up sequentially with one still folded over the receptacle. Surrey, England, UK
Common rue (Ruta graveolens) flower in UV light, pollen and nectar at base of petals fluorescing. Insect pollinators are able to see position of rewards on offer. Surrey, England, UK
Porcelain orchid (Chloraea magellanica) with osmophores on lip fluorescing in UV light. Chile. Controlled conditions, focus stacked. Sequence 2/2
Lily (Lilium lophophorum) flowers. Tepals fused at tips to protects floral parts from precipitation, slits allow access to pollinators. Mugecuo Scenic Area, Kangding, Sichuan Province, China. June
Leaf cutter ants (Atta sp) carrying piece of leaf, Costa Rica
Portrait of rescued and rehabilitated Grey-headed flying-foxes (Pteropus poliocephalus) that have just been transported to a soft-release enclosure within the Yarra Bend Park colony (Victoria)
Grey-headed Flying-foxes (Pteropus poliocephalus) at a colony hang together on a branch over the Yarra River (Melbourne). ÔÇ¿ These bats form part of a colony that was established here in early 2000s
Trailing fuschsia (Fuchsia procumbens) cultivated in alpine house, Surrey, England, UK. Native to New Zealand
Chinese alligator (Alligator sinensis) group from above, heads above water of Yangtze. Critically endangered species, individuals released from captive breeding programme, Anhui Province, China. 2009
Shoal of colorful yellow fish Panamic porkfish (Anisotremus taeniatus), Cabo Pulmo Marine National Park, Baja California Sur, Mexico
Shoal of Dog snapper, (Lutjanus novemfasciatus) in front of shoal of Big-eye jacks (Caranx sexfasciatus), Cabo Pulmo Marine National Park, Baja California Sur, Mexico
Great horned owl (Bubo virginianus) preening two chicks in nest in Saguaro (Carnegiea gigantea) cactus, Sonoran desert, Arizona, USA
Gentoo penguins (Pygoscelis papua) walking in line, returning to nesting area, Port Charcot, Antarctic Peninsula, Antarctica
Blacktip reef shark (Carcharhinus melanopterus) swims through schools of fish. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Killer whales (Orcinus orca) at the surface in an Arctic fjord in Spildra, northern Norway, with sea birds. Arctic Ocean
Several mimic goatfish (Mulliodichthys mimicus) hide within a school of Bluestripe snapper (Lutjanus kasmira) on a coral reef. North Ari Atoll, Maldives. Indian Ocean
School of White collar butterflyfish (Chaetodon collare) pack together above a coral reef. North Male Atoll, Maldives. Indian Ocean
Scarlet Macaws (Ara macao) drinking water inside an Amazonian claylick. Yasuni National Park, Orellana, Ecuador
Brent geese (Branta bernicla) flying with moon, Titchwell, Norfolk, UK, December
Common crane (Grus grus) flock in flight, Lac du Der, France, November
Flock of Pink footed geese (Anser brachyrhynchus) in flight at dusk, Norfolk, England, UK, October
People watching the Fissure eruption of Chico volcano into 9km diameter caldera. Sierra Negra volcano, Isabela Island, Galapagos, Ecuador. October 2005
Lesser horseshoe bats (Rhinolophus hipposideros) in magnesium mine, Shropshire, England, UK, April
King penguins (Aptenodytes patagonicus) walking in line on a windy beach. Sanders Island, Falklands. November
English oak (Quercus robur) ripening acorns, Dorset, UK September
Sulphur tuft mushrooms (Hypholoma fasciculare), Hampstead Heath, England, UK. September
Turnstones (Arenaria interpres) and Purple sandpipers (Calidris maritima) over wintering on the beaches of the Orkney Isles, Scotland, UK, April
Orca (Orcinus orca) surfacing, Shetland, Scotland, UK, August
Gannet (Morus bassanus) diving for fish, Noss, Shetland Islands, Scotland, UK, August
Group of Tissue moths (Triphosa dubitata) hibernating in a limestone cave. Peak District National Park, Derbyshire, UK. November
Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico
White-fronted goose (Anser albifrons) and Taiga bean goose (Anser fabalis rossicus) flocks in flight and on water. Latvia. April
Wirehaired daschund, Staffordshire bull terrier and Hungarian viszla in autumn colours, Wiltshire, UK
Black skimmers (Rynchops niger), three in non-breeding plumage on beach, Fort De Soto Park, St. Petersburg, Florida, USA. January. Digitally altered to remove distractions
White pelicans (Pelecanus erythrorhynchos) at Bear River Migratory Bird Refuge with an approaching storm in May, Utah, USA, May
Red-breasted Mergansers (Mergus serrator) male in foreground performing courtship display to female in background (left), Aurora, New York, USA, April
Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October
Wilsons Phalaropes (Phalaropus tricolor) flock flying over tufa formations at Mono Lake in late summer, with reflection, California, USA. July 2013
Avian feeding frenzy with a flock of White Pelicans (Pelecanus erythrorhynchos) swimming along a channel chasing school of fish
Mixed Blackbird flock, mostly Red-winged Blackbirds (Agelaius phoeniceus), in flight against backdrop of autumn foliage in November, Bosque Del Apache National Wildlife Refuge, New Mexico, USA
Snow buntings (Plectrophenax nivalis) flock in flight during a snowstorm, New York, USA, January
Snow Geese (Chen caerulescens) large flock in pool at Bosque Del Apache National Wildlife Refuge with setting full moon at sunrise in December, New Mexico, USA. December
Black-bellied whistling-ducks (Dendrocygna autumnalis), group perched amid Spanish moss on bald cypress tree, Lakeland, Florida, USA, January
European otter (Lutra lutra) group with one shaking off water, Kiskunsagi National Park, Hungary, October
Guanaco (Lama guanicoe) grazing on grassland plain, Patagonia, Chile. Novmeber
Llama (Lama glama) herd, Laguna colorada. Altiplano, Bolivia
Vicuna (Vicugna vicugna), five running across salt flat with mountain in background. Salar de Ascotan, Chile. September 2018