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RF - A vibrant Red Sea coral reef scene, with orange female Scalefin anthias fish (Pseudanthias squamipinnis) teeming over Fire coral (Millepora dichotoma)
Diver swimming with an aggregation of Cod (Gadus morhua) over a kelp forest. These cod were gathered in early spring off the north coast of Iceland to spawn. Thorshofn, Iceland. North Atlantic Ocean
Gemsbok (Oryx gazella) aerial view of herd on sand, Namib Desert, Namibia
Leaf cutter ants (Atta sp) carrying pieces of fern, reflected in water, Costa Rica
Bramble (Rubus plicatus) leaves and stem against white background. France, August
Blue and gold fusilier / Scissor tailed fusilier (Caesio caerulaurea) school swimming through shafts of sunlight, West Papua, Indonesia
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
Grants zebra (Equus burchelli boehmi) herd with large cloud above, Amboseli National Park, Kenya
Cape buffalo (Syncerus caffer) crossing the Chobe River, followed by swarm of flies, Chobe Game Reserve, Botswana, Africa
Gemsbok (Oryx gazella) female with two calves, Namibrand Reserve, Namib Desert, Namibia, Nambia
Caribbean flamingo (Phoenicopterus ruber) about to take off to roosting site, Ria Celestun Biosphere Reserve, Yucatan Peninsula Mexico, January
Portuguese Man-of-War (Physalia physalis) close up of tentacles, Sargasso Sea, Bermuda
Spotted eagle ray (Aetobatus narinari) school, Kagi kandhu, ; North Male Atoll, Maldives, Indian Ocean
Blue spotted trevally (Caranx bucculentus) hunting small fish in the reef, South Ari Atoll, Maldives, Indian Ocean
Common poppies (Papaver rhoeas) backlit in evening light, Polly / Porth Joke, Pentire, Newquay, Cornwall, UK, July
Trunks of giant sequoia trees (Sequoiadendron giganteum) Sequoia National Park, California, USA, November 2012
RF - American bisons (Bison bison) in Grand Teton National Park. winter. Wyoming, USA
Ring-tailed lemur (Lemur catta) female carrying two babies. Anjaha Community Conservation Site, near Ambalavao, Madagascar
Rothschild's giraffes (Giraffa camelopardalis rothschildi), feeding on leaves, Woburn Safari Park, UK, June, captive
Black howler monkeys (Alouatta caraya) male and two females calling from tree, captive, Apenheul Park, Netherlands
Spring crocus (Crocus vernus) flowering en masse on the Campo Imperatore, Abruzzo, Italy. April
Honeybee (Apis melifera) visiting Lavender (Lavendula angustifolia) in lavender fields, Valensole Plateau, Alpes Haute Provence, France, July
Black bear cubs (Ursus americanus) standing in a tree, Minnesota, USA, June
Thimble jellyfish (Linuche unguiculata) aggregation, My Paradise Reef, Cabilao Island, Bohol, Central Visayas, Philippines, Pacific Ocean
Three False clownfish (Amphiprion ocellaris) in Sea anemone (Heteractis magnifica) Lighthouse Reef, Cabilao Island, Bohol, Central Visayas, Philippines, Pacific Ocean
Atlantic salmon (Salmo salar) migrating for spawning in river, Gaspe Peninsula, Quebec, Canada, October
Three Strawberry poison frogs (Oophaga pumilio) on rock, Sarapiqui, Heredia, Costa Rica
Desert dwelling African elephants (Loxodonta africana) matriarch leading her family in the Hoanib River bed, Damaraland. Namibia
African elephants (Loxodonta africana) swimming across Chobe River, trunks raised above surface for breathing, Chobe National Park, border between Botswana and Namibia
Group of African elephants (Loxodonta africana) silhouetted at sunrise, Okavango Delta, Botswana
Spinner dolphins (Stenella longirostris) pod swimming over a shallow sandy lagoon in a coral reef. Sataya Reef, Fury Shoal, Egypt. Red Sea
Pod of Risso Dolphin (Grampus griseus) at surface, Baja California, Mexico
Large pod of Bottlenose dolphins (Tursiops truncatus) porpoising over waves during annual sardine run, Port St Johns, South Africa
Pod of Spinner dolphins (Stenella longirostris) swimming close to the surface. Sataya Reef, Fury Shoal, Marsa Alam, Egypt. Red Sea
Caribbean flamingo (Phoenicopterus ruber) flock flying in formation, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Whooper swans (Cygnus cygnus) group of three, camouflaged on frozen lake, Kussharo, Hokkaido Japan
Banded demoiselle (Calopteryx splendens) group of four on plant stems, River Leijgraaf, Nijmegen, the Netherlands, August 2013
Waterfowl at dawn with mist, including Great blue heron (Ardea herodias herodias), American avocet (Recurvirostra americana), Northern shoveler (Anas clypeata), Xochimilco wetlands, Mexico City
Oceanic manta ray (Mobula birostris) accompanied by three Black jacks (Caranx lugubris) silhouetted, Revillagigedo Islands, Mexico, Pacific Ocean
Caribbean reef sharks (Carcharhinus perezi) swimming near Eleuthera Island, Bahamas, North Atlantic Ocean
Zebras (Equus burchelli) herd during the Great Migration, Masai Mara National Park, Kenya. July
Mixed herd of Masai giraffe (Giraffa camelopardalis) walking across short grass plains in the shadow of Oldonyo Lengai, Ngorongoro Conservation Area, Tanzania
Japanese / Red-crowned crane (Grus japonicus) three in flight, Hokkaido Japan February
A flock of Starlings (Sturnus vulgaris) flying in front of the Blackpool Tower, England, UK, September 2010
Caribbean flamingo (Phoenicopterus ruber) two walking, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, September. Bookplate
Two Great egrets (Ardea alba) standing opposite each other with Grey heron (Ardea cinerea) in between. Lake Csaj, Pusztaszer, Hungary, January
Cheetah (Acinonyx jubatus) female, with three cubs aged approx. 8 months, sitting side by side at dawn, Ndutu area, Serengeti / Ngorongoro Conservation Area, Tanzania