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Sergeant major (Abudefduf sp) schooling in mangroves of Isabela Island, Galapagos
Red deer (Cervus elaphus) herd during rut in morning light. Derbyshire, England, UK. October
Marsh fritillary (Euphydryas aurinia), group feeding on Marsh thistle (Cirsium palustre). Lincolnshire, England, UK
Green-crowned brilliant (Heliodoxa jacula), three feeding on Lobster-claw (Heliconia sp) flower in rain. Costa Rica
Black-headed gull (Chroicocephalus ridibundus) flock flying over a frozen lake in falling snow, Wiltshire, UK, March
Greater flamingos (Phoenicopterus roseus) flock, Sado Estuary, Portugal. January
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Pack of wolves (Canis lupus). Yellowstone National Park, Wyoming, USA. January
Thomsons gazelle (Eudorcas thomsonii) herd. Serengeti National Park, Serengeti, Tanzania
Group of Swan geese (Anser cygnoides) on grassland, Inner Mongolia, China
Piraputanga, (Brycon hilarii) reflected on the water surface, Aquario Natural, Bonito, Mato Grosso do Sul, Brazil
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Bohar snapper (Lutjanus bohar) school at Ras Mohammed, Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea
Masai giraffe (Giraffa camelopardalis tippelskirchi) male, females and calf standing close together, Masai Mara Game Reserve, Kenya
Thomsons gazelle (Eudorcas thomsonii) male and two females standing in rainstorm, Masai Mara Game Reserve, Kenya
Whale Shark (Rhincodon typus) and Golden trevally (Gnathanodon speciosus) Cenderawasih Bay, West Papua, Indonesia
Killer whale (Orcinus orca) hunting inside herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Cockroaches (Periplaneta americana) on wall of Gomantong Cave. Sabah, Borneo. Malaysia
Galapagos racer snakes (Pseudalsophis biserialis) group alert watching for prey on beach, Galapagos, June. Endangered species. Taken on location for BBC tv Blue Planet 2 series
Flock of Mallard ducks (Anas platyrhynchos) taking off, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. December
Blackcap (Sylvia atricapilla) female and two males huddling together for warmth before going to sleep, Finland, May
Common redpoll (Acanthis flammea), flock feeding, Finland, January
Southern Rockhopper penguin (Eudyptes chrysocome) colony, Kidney Island, Falkland Islands, October. Vulnerable species
Gentoo penguin (Pygoscelis papua) walking back to the colony, Sea Lion Island, Falkland Islands, October
Fallow deer (Dama dama) at night, Gyulaj, Hungary Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Meerkats (Suricata suricatta), Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Eastern Ring-tailed Vontsira (Galidia elegans elegans) (endemic family Eupleridae) foraging on forest floor. Marojejy National Park, north east Madagascar
Freshwater isopods (Acanthogammarus victorii) feeding on seabed, Lake Baikal, Siberia, Russia
Indian wild ass (Equus hemionus khur), group standing together, Little Rann of Kutch, Gujarat, India
Starling (Sturnus vulgaris), eating fruit of Japanese Persimmon (Diospyros kaki), Provence, France
Southern elephant seal (Mirounga leonina), fight between two males, Saint Andrew, South Georgia
Series of Bald eagle (Haliaeetus leucocephalus) in flight, Homer, Alaska, USA, March. Digital composite
Puna / James flamingo (Phoenicoparrus jamesi) flock on Laguna Colorado, Reserva Eduardo Avaroa, Altiplano, Bolivia
Great Frigatebird (Fregata minor) arriving at nest with chick, San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
European Chubb (Squalius / Leuciscus cephalus) in cave, Plitvice Lakes National Park, Croatia. November
School of Munks devil ray (Mobula munkiana), feeding on plankton at night, photographed on a long exposure, Espiritu Santo Island, Sea of Cortez, Baja California, Mexico, East Pacific Ocean
Grey wolf (Canis lupus) mother and two month old cubs, pups begging for food by licking mothers mouth, captive
Red Deer (Cervus elaphus) herd in forest in snow, Cairngorms National Park, Scotland, UK. December
Polar bear (Ursus maritimus) sow with two cubs walking past a pile of Bowhead whale (Balaena mysticetus) bones, Kaktovik, Barter Island, 1002 Area, Arctic National Wildlife Refuge, North Slope
Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Pink-footed goose (Anser brachyrhynchus) flock flying in mist, Cromarty Firth, Highlands, Scotland, UK April
Curlew (Numenius arquata) roosting on saltmarsh, Cromarty Firth, Scotland, UK April
Chilean devil rays / Mobulas (Mobula tarapacana) offshore Santa Maria, Azores, Portugal, Atlantic Ocean
Greater flamingos (Phoenicopterus roseus) flock reflected, feeding in water, Pont de Gau Ornithological Park, Camargue, France, March
Leaf-cutter ants (Atta cephalotes) carrying pieces of leaf that they have harvested back to their underground fungus garden in their nest, Osa Peninsula, Costa Rica
Plankton at night, Palau, Philippine Sea
Army ants (Eciton sp. ) Costa Rica. February 2015Army ants (Eciton sp.) Costa Rica. February 2015
Alpine ibex (Capra ibex) group posing on a rocky mountain side on misty day, Gran Paradiso National Park, Italy. November
