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Chicken wire coated in hoar frost. Peak District National Park, Derbyshire, UK, December 2012
Mixed species woodland in autumn, including Silver birch trees (Betula pendula), Peak District National Park, England, UK, October
Ice pattern, Merrivale, Dartmoor National Park, Devon, England, UK, February
Portrait of a European grey wolf (Canis lupus), captive, Norway, February
Cracks in the ice of Lake Baikal, Siberia, Russia, March
Wood patterns in Lodgepole pine (Pinus contorta) Yellowstone National Park, Wyoming, USA, January
Portrait of an Arctic fox (Vulpes lagopus), captive, Norway, February
Antelope Canyon, a slot canyon with eroded sandstone patterns, Navajo Tribal Park, Arizona, USA, December 2012
Mist in the cloud forest at Sombom Ridge, Saruwaged Range, Huon Peninsula, Papua New Guinea
Leopard (Panthera pardus) portrait, Samburu National Reserve, Kenya
Coastal brown bear (Ursus arctos) resting, Lake Clarke National Park, Alaska, September
Cloud formations lit by low sun. Brechin, Angus, Scotland, December 2012
White-cheeked pintail (Anas bahamensis galapagensis) resting, Santa Cruz Island, Galapagos
Peruvian screech-owl (Megascops roboratus) half face close up portrait, Macara, Loja, Ecuador
RF - Moths attracted to light trap in rainforest, on white background, image composite montage. Cordillera de Talamanca mountain range, Caribbean Slopes, Costa Rica
In-camera motion blur image of a female Yellow Seahorse (Hippocampus kuda). Lembeh Strait, Sulawesi, Indonesia, Molluca Sea, March
Close up detail of soft coral (Dendronephthya sp. ) Maldives, Indian OceanClose up detail of soft coral (Dendronephthya sp.) Maldives, Indian Ocean
Maple leaves backlit on lightbox, Broxwater, Cornwall, UK. November 2013
Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October
Snakelocks Anemone (Anemonia viridis), green form in a rockpool, Devon, UK. June
Green aurora borealis light in night sky, Senja, Norway, February 2012
RF- Bluebells (Hyacinthoides non-scripta / Endymion scriptum) flowering in Broad buckler fern (Dryopteris dilatata), Hallerbos, Belgium. April
Arizona mountain kingsnakes (Lampropeltis pyromelana) Found in Chiricahua mountains, SE Arirona. Controlled conditions
Wildebeest (Connochaetes taurinus) migration, herd crossing the Mara river, Masai-Mara Game Reserve, Kenya
Shoaling glassfish (Parapriacanthus sp. ) shoal, Maldives, Indian OceanShoaling glassfish (Parapriacanthus sp.) shoal, Maldives, Indian Ocean
Lobelia (Lobelia deckenii) close up of leaves, alpine species of lobelia that lives on Kilimanjaro, Tanzania
Monarch butterfly (Danaus plexippus) hibernating, Mariposa Monarca Special Biosphere Reserve, central Mexico, January
Flock of Greater flamingo (Phoenicopterus ruber) in flight, Camargue, Southern France, November 2006
Aerial view of salt pans coloured red by cyanobacteria, Lake Natron, Rift Valley, Tanzania, August 2009
Aerial view of patterns created by agricultural work on land, Los Monegros, Zaragoza Province, Aragon, Spain, July 2011
Highland bull (Bos taurus) face close up in freezing fog, Berwickshire Scotland, December
Aggregation / bloom / swarm of Moon Jellyfish (Aurelia aurita) in the clear waters of Loch Na Keal, off the Isle of Mull, north west Scotland, UK, June, Photographed through the surface
Indian elephant (Elephas maximus) close up of skin and tail, captive
Red river hog (Potamochoerus porcus) close up of head, captive
Red fox (Vulpes vulpes) head portrait, The Netherlands, March
Tulips (Tulipa) Synaeda Orange, cultivated, Schwerin, Germany
Crowned crane (Balearica regulorum) close-up of feathers, captive
Greater flamingo (Phoenicopterus ruber) close-up of back, captive
Male Indian peafowl (Pavo cristatus) displaying, captive
Black swan (Cygnus atratus) captive
Agave (Agave sp. ) leaf close-up detail. Maui, Hawaii, FebruaryAgave (Agave sp.) leaf close-up detail. Maui, Hawaii, February
Composite image of flying insects visiting blacklight, Texas, USA, September
Southern elephant seal (Mirounga leonina) close up of sleeping individual, South Georgia. Taken on location for BBC Frozen Planet series, 2008
Looking up through the canopy of Fan Palms (Licuala ramsayi) in rainforest, Daintree National Park, North Queensland, Australia
Termites (Isoptera) flying upwards towards a street lamp, India
Wild / feral Dulmen ponies (Equus caballus) close up of herd of mares and foals running on the Duke of Croys estate, Meerfelder Bruch, North Rhine-Westphalia, Germany, May 2011
Jackdaws (Corvus monedula) abstract view of flock in flight, Derbyshire, UK, December. Non-ex
African elephants (Loxodonta africana) bulls crowd into an emptying waterhole to drink, Savuti, Botswana. Taken on location for BBC Planet Earth series, 2005
