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School of Bluespine unicornfish (Naso unicornis) gathered during the summer breeding season. Yolanda Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Close-up portrait of a European grey wolf (Canis lupus), captive, Norway, February
Keel billed toucan (Ramphastos sulfuratus), captive
Brown Hare (Lepus europaeus) close-up of fur and eye. Derbyshire, UK, March
Lesser flamingos (Phoeniconaias minor) flock in Lake Bogoria, Rift valley, Kenya
Violet bramble rust (Phragmidium violaceum) causing red colouration on Bramble leaf (Rubus fruticosus aggregate). Peak District National Park, Derbyshire, UK. October
Eyes of Margined sole fish (Brachirus heterolapis) on its camouflaged body. Saonek Island, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean. Dampier Strait
A Mountainous star coral (Montastraea faveolata) spawning at night, with gamete bundle being released from a coral polyp, East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Purple short tentacle mushroom coral (Fungia sp. ) close up of polyps. Lembeh StraitPurple short tentacle mushroom coral (Fungia sp.) close up of polyps. Lembeh Strait, North Sulawesi, Indonesia
Lavender (Lavandula angustifolia) fields, Valensole Plateau, Alpes Haute Provence, France, June
Gorgonian / Fan coral (Solenocaulon akalyx), close up, West Papua, Indonesia
RF- Chanterelle mushroom (Cantharellus cibarius), gill detail
Coral clam (Pedum spondyloideum) close up, Maldives, Indian Ocean
Fallen leaves in pond with aquatic plants, Denmark, Europe, September
Great Green Macaw (Ara ambiguus) close up of feathers, El Manantial Macaw Sanctuary, Costa Rica. Captive
Great yellow gentian (Gentiana lutea) leaves. Haute-Savoie, Massif des Bauges, French Alps, May
RF- Boulder star coral (Montastrea annularis) showing fluorescent green coloration, photographed under blue light at night, to excite fluorescent pigments in the coral, East End, Grand Cayman
Brain coral (Diploria strigosa) photographed under blue light to excite the fluorescence, Little Cayman, Cayman Islands, British West Indies, Caribbean Sea
Eastern green mamba {Dendroaspis angusticeps} skin detail, captive, from East Africa
Bigeye trevally or jack (Caranx sexfasciatus) shoal, Sipadan, Malaysia
RF- Close up of Leaves of Red Oak (Quercus rubra) in autumn. October
Wildebeest (Connochaetes taurinus) crossing the Mara river, Masai-Mara Game Reserve, Kenya
Detail of sponge (Chalinula nematifera) Maldives, Indian Ocean
Broken ice abstract, Jokulsarlon glacier lagoon. Southern Iceland, Iceland, Europe, March 2011
Detail in frozen waterfall. Selijalandfoss waterfall, Southern Iceland, Europe, March 2011
Blue-and-Yellow Macaw (Ara ararauna) close up of feathers. Captive, Brazil, South America
Aerial view saltmarshes of Bahia de Cadiz Natural Park, Huelva, Andalucia, Spain, March 2008
RF- Secretary blenny (Acanthemblemaria maria) peering from hole in massive Starlet coral (Siderastrea siderea). East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Abstract of reeds in front of lake with reflections, Plitvice Lakes National Park, Lika, Croatia, Europe, October 2011
Leaves floating on water with colourful reflections, Plitvice Lakes National Park, Lika, Croatia, Europe, October 2011
Gloucester old spot piglets (Sus scrofa domestica) suckling at the Cotswold Farm Show, Cirencester, UK, July 2008
Rothschilds giraffe (Giraffa camelopardalis rothschildi) close up of young calf skin pattern, captive
Grevy's zebra (Equus grevyi) close up, captive
Bigeye Scad (Selarcrumenophthalmus). Misool, Raja Ampat, West Papua, Indonesia
Aerial view of the rivers and saltmarshes of the Bahia / Bay de Cadiz Natural Park, Andalucia, Spain, March 2008
Looking up at Beech wood canopy (Fagus sylvatica) in autumn, Peak District National Park, Derbyshire, UK, November
River patterns from melting glacier. Olfusa river, Southwest Iceland, July 2009
Aerial view of Lake Natron, Rift Valley, Tanzania, Africa, August 2009
Pink abalone (Haliotis corrugata) shell detail, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, January
Close up of coiled tail of Parsons chameleon {Calumma parsonii} showing scales and strong green colouration. Tropical rainforest, Masoala Peninsula National Park, north east Madagascar
Detail in ice pattern formations, the Netherlands
Wild flower meadow in city centre, Sheffield, Yorkshire, UK, August 2009
King Penguin (Aptenodytes patagonicus) adult standing amongst colony of chicks, Gold Harbor, South Georgia, South Atlantic Islands
Koala (Phascolarctos cinereus) mother with joey aged eight months, Queensland, Australia, captive
Head portrait of Highland cow, Scotland, with tiny frond of bracken at corner of mouth, UK
Female African lion (Panthera leo) face portrait, Etosha National Park, Harare Province, Namibia
Close-up of an eye of a Sumatran tiger (Panthera tigris sumatrae), captive, occurs in Sumatra, Indonesia
Portrait of Siberian Husky dog face used as sled dogs inside Riisitunturi National Park, Lapland, Finland