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Pink anemonefish (Amphiprion perideraion) in Purple magnificent sea anemone (Heteractis magnifica) Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Reticulate stargazer (Uranoscopus sp. ) emerging from the sandReticulate stargazer (Uranoscopus sp.) emerging from the sand. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
RF- Wide angle portrait of Hairy frogfish (Antennarius striatus) waiting to ambush prey on black sand slope. Lembeh Strait, North Sulawesi, Indonesia. Molucca Sea
Portrait of Paddle-flap scorpionfish (Rhinopias eschmeyeri) on rubble slope. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea
Maldive anemonefish (Amphiprion nigripes) shoal swimming in the current flowing over a Magnificent sea anemone (Heteractis magnifica). Baa Atoll, Maldives. Indian Ocean
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon, on coral reef under stormy seas. St Johns Reef. Egypt. Red Sea
Butterfish (Pholis gunnellus) portrait. Gardur, south west Iceland. North Atlantic Ocean
Red Irish lord (Hemilepidotus hemilepidotus) hides on the sebed, with Quillback rockfish (Sebastes maliger) and copper rockfish (Sebastes caurinus) behind in the bull kelp forest
Blue sharks (Prionace glauca) cruise beneath the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August
Roughhead blenny (Acanthemblemaria aspera) male portrait. This species is very small, just 20mm long and 1-2mm deep, East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Secretary blenny (Acanthemblemaria maria) making a home in a deserved worm tube of a great star coral (Montastraea cavernosa), East End, Grand Cayman, Cayman Islands, British West Indies
Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Tassled anglerfish (Rhycherus filamentosus) is almost perfectly camouflaged as it hides amongst marine life, waiting to ambush prey attracted to worm life lure
Spotted handfish (Brachionichthys hirsutus) portrait, Derwent River estuary, Hobart, Tasmania, Australia
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
Whale shark (Rhincodon typus) silhouette, feeding on floating fish eggs (not visible) just below the surface in calm weather. Note the bow wave ahead of the shark
RF - Long exposure of a school of yellowtail surgeonfish (Prionurus punctatus), these fish school to help them access the best feeding areas, which are controlled by aggressive damselfish
Whitetip shark (Carcharhinus longimanus), accompanied by Pilotfish (Naucrates ductor), swims towards the camera as it cruises beneath the surface of the Red Sea, close to Little Brother Island
Lionfish (Pterois volitans) portrait, invasive species in the Caribbean. East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Portrait of a male Squarespot anthias (Pseudanthias pleurotaenia). Sangeang Island; Sumbawa, Indonesia. Flores Sea
School of Blacksmith fish (Chromis punctipinnis) swim through a giant kelp (Macrocystis pyrifera) forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Hairy frogfish (Antennarius striatus) on a sandy seabed. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Bumphead parrotfish (Bolbometopon muricatum) gather at dawn on a shallow coral reef. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea. Vulnerable species
Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean
Orange-finned anemonefish (Amphiprion chrysopterus) guarding their recently spawned red eggs on a rock next to the host anemone. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean
Puffin (Fratercula arctica) portrait with beak full of sand eels, Farne Islands, Northumberland, UK, June
A digital composite of a Pygmy Seahorse (Hippocampus bargibanti) swimming over a seafan (Muricella sp.). Pygmy seahorses are small, most less than 15mm in total length
Giant Pacific Manta Ray (Manta birostris) portrait with Remora fish (Echeneididae) Roca Partida, Revillagigedos Islands, Mexico. East Pacific Ocean. February
Draughtsboard Shark / Australian Swellshark (Cephaloscyllium laticeps) swims through a giant kelp forest (Macrocystis pyrifera)
A large Great hammerhead shark (Sphyrna mokarran), about 4m in length, cruises overhead. Bahamas, Tropical West Atlantic Ocean
Pink anemonefish (Amphiprion perideraion) in a purple Magnificent sea anemone (Heteractis magnifica). Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Striped triplefin (Helcogramma striata) on hard coral, Tubataha Reef, Philippines
Spinyhead blenny (Acanthemblemaria spinosa) in hard coral, Netherlands Antilles, Bonaire, Caribbean
Emperor angelfish (Pomacanthus imperator) swimming past soft corals (Dendronephthya sp]), Red Sea, Egypt. January
A Large / Pot Bellied Seahorse (Hippocampus abdominalis) perched in nets. Manly, Sydney, New South Wales, Australia, March
DELETE Striped marlin (Kajikia audax) feeding on bait balls, Eastern Pacific, Thetis Bank, Baja California, Mexico
School of Blackfin barracuda (Sphyraena qenie), Kimbe Bay, West New Britain, Papua New Guinea
Yellow and blueback fusilier (Caesio teres) schooling over coral reef, West Papua, Indonesia
RF - Young lionfish (Pterois volitans) swimming near surface hunting silversides, at dusk. Beacon Rock, Sha ab Mahmood, Sinai, Egypt. Red Sea
RF- Elongate surgeonfish (Acanthurus mata) school. Indonesia
Napoleon wrasse (Cheilinus undulatus) portrait. Red Sea, Egypt. October
Yellow and blueback fusilier (Caesio teres) shoal, Similan Islands, Andaman Sea, Thailand
Bigeye trevally or jack (Caranx sexfasciatus) shoal, Sipadan, Malaysia
Atlantic bluefin tuna (Thunnus thynnus) within tuna farm, containing around 1000 per net. Saint Pauls Bay, Malta. Mediterranean Sea
Close up of pattern of adult Emperor angelfish (Pomacanthus imperator) Tulamben, Bali, Indonesia. Java Sea
Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
