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Ratfish / Ghost shark (Chimaera monstrosa) on fjord bed, Trondheimsfjorden, Norway, November 2006
Atlantic bluefin tuna (Thunnus thynnus) portrait, captive, Malta, Mediteranean, May 2009 WWE BOOK
Monkfish / Allmouth (Lophius piscotorius) close-up of face, Lofoten, Norway, November 2008
Anglerfish (Lophius piscatorius) Saltstraumen, Bod, Norway, October 2008
Catch of Atlantic mackerel (Scomber scombrus) in fish separator on board Shetland pelagic trawler Charisma, Shetland Isles, Scotland, UK, October 2012
Fraserburgh pelagic trawler Forever Grateful fishing for Atlantic mackerel (Scomber scombrus) close to Papa Stour, Shetland Isles, Scotland, UK, October 2012
Male Black faced blenny (Tripterygion delaisi), part of a sequence showing colour change, Swanage, Dorset, England, UK, May. 2/4
Shanny (Lipophrys pholis) in a crevice in a harbour wall, Bovisand Harbour, Plymouth, Devon, England, UK, July
John dory (Zeus faber), Porthkerris Cove, Cornwall, England, UK, June
Basking shark (Cetorhinus maximus) feeding at the surface on plankton, Cairns of Coll, Inner Hebrides, Scotland, UK, June
Tompot blenny (Parablennius gattorugine) beneath Swanage Pier, Dorset, UK. Photographed during May. English Channel
Brown trout (Salmo trutta) in turbulent water at a weir, River Ettick, Selkirkshire, Scotland, UK, October
Atlantic wolf fish (Anarhichas lupus) in a rock crevice, St Abbs (St Abbs and Eyemouth Voluntary Marine Reserve), Berwickshire, Scotland, UK, August 2011
RF- Rear view of Basking shark (Cetorhinus maximus) feeding on plankton, visible as white dots on water surface near island of Coll, Inner Hebrides, Scotland, UK. June
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea in the late afternoon, close to Little Brother Island
Snorkeller with shoal of Piraputanga (Brycon hilarii), Rio Olha d Agua at Cabeceira do Prata, Bonito, Mato Grosso do Sul, Brazil
Great Fen / Raft spider (Dolomedes plantarius), adult female eating an invasive species of fish, Western mosquitofish, (Gambusia affinis), Alessandria, Italy, August
Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July
Kingfisher (Alcedo atthis) male passing fish to female spring courtship behaviour, Hertfordshire, England, UK, March. Sequence 4 of 6
Portrait of Blacktip grouper (Epinephelus fasciatus) resting in Barrel sponge (Xestospongia testudinaria) Mabul Island, Sabah, Borneo, Malaysia. Sulawesi Sea
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by a group of Pilotfish (Naucrates ductor). Rocky Island, Egypt. Red Sea
School of large Atlantic bluefin tuna (Thunnus thynnus) captive in growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
Large male Napoleon wrasse (Cheilinus undulatus) yawning. Ras Mohammed, Sinai, Egypt. Red Sea
School of Longnose parrotfish (Hipposcarus harid) swarming over reef. Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Basking shark (Cetorhinus maximus) feeding on plankton in the surface waters around the island of Coll, Inner Hebrides, Scotland, UK, June. 2020VISION Book Plate
Southern stingray (Hypanus americanus) female swimming over a shallow sand bank, under morning sun, North Sound, Grand Cayman, Cayman Islands. British West Indies, Caribbean Sea
RF- School of Blackfin barracuda (Sphyraena qenie) forming circle in open water at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Black rockfish (Sebastes melanops) in bull kelp (Nereocystis luetkeana), Browning Pass, Port Hardy, Vancouver Island, British Columbia, Canada
Klunzingers wrasse (Thalassoma klunzingeri) swimming over the edge of fringing reef where the water depth drops instantly from few feet to hundreds of meters. Ras Mohammed Marine Park, Sinai, Egypt
Male Yellow-headed jawfish (Opistognathus aurifrons) blows out the clutch of eggs he is incubating in his mouth to oxygenate them. East End, Grand Cayman, Cayman Islands, British West Indies
A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Pair of Sockeye salmon (Oncorhynchus nerka) on their redd in a shallow stream. Female in front of male. Huihil Creek, Adams River, British Colombia, Canada, October
Portrait of Male Sockeye salmon (Oncorhynchus nerka) showing teeth. Adams River, British Columbia, Canada, October
Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
RF - A vibrant Red Sea coral reef scene, with orange female Scalefin anthias fish (Pseudanthias squamipinnis) teeming over Fire coral (Millepora dichotoma)
RF - Close up detail of the eye and skin pattern of a Giant hawkfish (Cirrhitus rivulatus). Socorro Island, Pacific Ocean
Diver swimming with an aggregation of Cod (Gadus morhua) over a kelp forest. These cod were gathered in early spring off the north coast of Iceland to spawn. Thorshofn, Iceland. North Atlantic Ocean
Yellowhead jawfish (Opistognathus aurifrons) spitting out sand as it digs its burrow. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
A Leafy Seadragon (Phycodurus eques), photographed from below. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia. Australia
Male Great white shark (Carcharodon carcharias) with sunrays, Guadalupe Island, Mexico, Pacific Ocean
RF - Banggai cardinalfish (Pterapogon kauderni). Lembeh Strait, North Sulawesi, Indonesia
Citron (lemon coral) goby (Gobiodon citrinus) Red Sea
Two Common butterfly fish (Chaetodon chrysurus) Red Sea, Egypt
Striped marlin (Kajikia audax) feeding on bait balls, Eastern Pacific, Thetis Bank, Baja California, Mexico
Blue and gold fusilier / Scissor tailed fusilier (Caesio caerulaurea) school swimming through shafts of sunlight, West Papua, Indonesia
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Blue triggerfish (Pseudobalistes fuscus) Egypt, Red Sea