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European chub (Squalius / Leuciscus cephalus) and Bream (Abramis brama) shoal in calm part of the Rhine, Schaffhausen, Switzerland, February 2009
Atlantic salmon (Salmo salar) migrating upstream to spawn, Umelven, Sweden, July 2009
Anglerfish (Lophius piscatorius) portrait, Saltstraumen, Bod, Norway, October 2008. WWE INDOOR EXHIBITION
Shrimp (Lebbeus polaris) eyeballing an Atlantic wolffish (Anarhichas lupus) Saltstraumen, Bod, Norway, October 2008
Anglerfish (Lophius piscatorius) underside, Saltstraumen, Bod, Norway, October 2008
Juvenile Atlantic wolffish (Anarhichas lupus) Saltstraumen, Bod, Norway, October 2008
Atlantic cod (Gadus morhua) portrait, Saltstraumen, Bod, Norway, October 2008
Coalfish / Saithe (Pollachius virens) Saltstraumen, Bod, Norway, October 2008
Atlantic cod (Gadus morhua) Saltstraumen, Bod, Norway, October 2008
Atlantic wolffish (Anarhichas lupus) in hole between rocks, Saltstraumen, Bod, Norway, October 2008
Painted comber (Serranus scriba) Turtle Rock, Passage du Cavallo, Lavezzi Archipelago, Corsica, France, September 2008
Dusky grouper (Epinephelus marginatus) by seagrass, Cala di Grecu, Lavezzi Islands, Corsica, France, September 2008
Osprey (pandion haliaetus) with fish prey, Cairngorms National Park, Scotland, UK, May. Did you know? Ospreys are found on every continent except Antarctica
Osprey (Pandion haliaetus) fishing, Cairngorms National Park, Scotland, UK, July
Atlantic mackerel (Scomber scombrus) in the net of Shetland pelagic trawler Charisma, Shetland Isles, Scotland, UK, October 2012
Puffin (Fratercula arctica) with beak full of sandeels, Farne Islands, Northumberland, England, UK, July
Dorsal fin of a Basking shark (Cetorhinus maximus) at the surface, Cairns of Coll, Inner Hebrides, Scotland. UK, June
Ballan wrasse (Labrus bergylta), Porthkerris cove, Cornwall, England, UK, June
Puffin (Fratercula arctica) with sand eels in beak, Farne Islands, Northumberland, June
Captured Atlantic salmon (Salmo salar) awaiting tagging by Tweed Foundation staff monitoring fish movement, density and biology, River Tweed, Berwickshire, Scotland, UK, September
Atlantic salmon (Salmo salar) leaping up the cauld at Philphaugh Salmon Viewing Centre near Selkirk, where members of the public can view the spectacle, Selkirkshire, Scotland, UK, October 2011
Brown trout (Salmo trutta), Ennerdale Valley, Lake District NP, Cumbria, England, UK, November 2011
Arctic charr (Salvelinus alpinus) males showing breeding colours, in a river where they have come to spawn, Ennerdale Valley, Lake District NP, Cumbria, England, UK, November 2011
Yellow / spiny seahorse (Hippocampus guttulatus) female sheltering is meadow of common Eelgrass (Zostera marina) Studland Bay, Dorset, UK, August
Rear view of Basking shark (Cetorhinus maximus) feeding on plankton in the surface waters around the island of Coll, Inner Hebrides, Scotland, UK, June
Basking shark (Cetorhinus maximus) disappears back into the blue in the surface waters around the island of Coll, Inner Hebrides, Scotland, UK, June
Lesser spotted catshark / Dogfish shark (Scyliorhinus canicula) hiding amongst seaweeds, Babbacombe Bay, Devon, UK, April
Brown Trout (Salmo trutta) in disused quarry, Jackdaw Quarry, Capernwray, Carnforth, Lancashire, UK, August
Detail of the pectoral fin of Roundhead parrotfish (Scarus viridifucatus) coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea
Whale shark (Rhincodon typus) with shoal of anchovies staying near to avoid predation. Los Roques, Venezuela. Caribbean
Harlequin sweetlips (Plectorhinchus chaetodonoides) with cleaning Bluestreak cleaner wrasse (Labroides dimidiatus) and Bicolour cleaner wrasse (Labroides bicolor) South Atoll, Tubbataha Atolls
Sockeye Salmon (Oncorhynchus nerka) travelling upstream on spawning migration, Adams river, British Columbia, Canada, October Taken for the Freshwater Project
School of Bluespine unicornfish (Naso unicornis) gathered during the summer breeding season. Yolanda Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Golf-ball sized Painted frogfish (Antennarius pictus) waiting to ambush prey disguised as an orange sponge. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Close up portrait of female Lionfish (Pterois volitans) in the late afternoon, when Lionfish are more active. The Alternatives, Sha ab Mahmood, Sinai, Egypt. Red Sea
Male Yellow boxfish (Ostracion cubicus) displaying, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
School of young European perch (Perca fluviatilis) in Altausseer lake, Austria, July
Great Blue shark (Prionace glauca) viewed from just below the surface, Pico Island, Azores, Portugal, Atlantic Ocean
Whale shark (Rhincodon typus) mouth open filter feeding at the surface, Quintana Roo, Yucatan Peninsular, Mexico, Caribbean Sea
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Freshwater Dorado (Salminus brasiliensis) Recanto Ecologico, Rio da Prata, Bonito, Mato Grosso do Sul, Brazil
A signal blenny (Emblemaria hypacanthus) displays its showy dorsal fin as it extends out of its tube home. La Paz, Mexico. Sea Of Cortez, East Pacific Ocean
Osprey (Pandion haliaetus) catching trout, Rothiemurchus estate, Cairngorms, Scotland, UK, July
Puffin (Fratercula arctica) in flight, with beak full of sand eels. Fair Island, Shetland Islands, Scotland, July
Eyes of Margined sole fish (Brachirus heterolapis) on its camouflaged body. Saonek Island, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean. Dampier Strait
Pink anemonefish (Amphiprion perideraion) in Purple magnificent sea anemone (Heteractis magnifica) Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Reticulate stargazer (Uranoscopus sp. ) emerging from the sandReticulate stargazer (Uranoscopus sp.) emerging from the sand. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
RF- Wide angle portrait of Hairy frogfish (Antennarius striatus) waiting to ambush prey on black sand slope. Lembeh Strait, North Sulawesi, Indonesia. Molucca Sea