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Red-footed booby (Sula sula) perched on branch, Genovesa Island, Galapagos Islands, Ecuador
Shoal of Mediterranean Fairy basslet (Anthias anthias) swimming between Red gorgonian (Paramuricea clavata) that have colonised the crossbars of a sunken electricity pylon that fell into the sea
Jellyfish (Thysanostoma sp. ) juvenile, drifting near the surface, Hawaii, Pacific OceanJellyfish (Thysanostoma sp.) juvenile, drifting near the surface, Hawaii, Pacific Ocean
Two Little bee-eaters (Merops pusillus) perched side by side on branch, one with insect in beak, Allahein River, The Gambia
Male Beautiful sunbird (Cinnyris Pulchellus) courtship display, Allahein River, The Gambia
Southern masked weaver (Ploceus velatus) building nest hanging from tree branch, Kgalagadi Transfrontier Park, South Africa
Beech (Fagus sp. ) trees in autumn, Saja Natural Park, Cantabria, Spain. NovemberBeech (Fagus sp.) trees in autumn, Saja Natural Park, Cantabria, Spain. November
Male Emperor angelfish (Pomacanthus imperator) swimming over garden of Fire corals (Millepora sp)Male Emperor angelfish (Pomacanthus imperator) swimming over garden of Fire corals (Millepora sp.) in morning sunshine, Ras Mohammed National Park, Sinai, Egypt, Red Sea
Emperor angelfish (Pomacanthus imperator) swimming in front of Fire corals (Millepora dichotoma) and soft corals (Dendronephthya sp.) and (Scleronephthya sp)
Powderblue surgeonfish (Acanthurus leucosternon) school swimming over a coral reef, Laamu Atoll, Maldives, Indian Ocean
Colourful wildflowers growing in arable farmland in summer, Ballinlaggan, Scotland, UK. July
Eyes of Red Irish lord (Hemilepidotus hemilepidotus) in Browning Pass, Vancouver Island, British Columbia, Canada. North East Pacific Ocean. July
Goldfinch (Carduelis carduelis) perched on twig in garden hedge in winter, Scottish Borders, Scotland, December
Fragrant Orchid (Gymnadenia conopsea) flowering, RSPB Insh Marshes, Cairngorms National Park, Scotland, UK. June
Raccoon butterflyfish (Chaetodon lunula), endemic Saddle wrasse (Thalassoma duperrey) and Sea urchin (Diadema sp)Raccoon butterflyfish (Chaetodon lunula), endemic Saddle wrasse (Thalassoma duperrey) and Sea urchin (Diadema sp.) on reef, Lanai, Hawaii, Pacific Ocean
Male Kingfisher (Alcedo atthis) sitting on eggs for his brooding period in an artificial nest, Italy
Diverse array of algae species including Sargassum weed (Sargassum horneri), Kelp (Laminariales), Sea lettuce (Ulva lactuca) on shallow reef with juvenile Pollock (Pollachius pollachius)
Festival clown in costume, Gom Kora Festival, Tashgangregion, eastern Bhutan 2001
Domani rock prayers outside Kurjey monastery, Bumthang, Central Bhutan 2001
Bower of Vogelkop Bowerbird (Amblyornis inornata) with bright decorations. Nov 2004
Alder (Alnus glutinosa) tree silhouetted by a display of Aurora borealis in the night sky, Cairngorms National Park, Scotland, UK. February
American Purple gallinule (Porphyrio martinica) balancing on a lily pad, Everglades National Park, Florida, USA
Close up of the tube feet of a Purple velvet seastar (Leiaster leachi), extending from the protective crevice in one of its arms, Maui, Hawaii, Pacific Ocean
Scalefin anthias / Scalefin basslet (Pseudanthias squamipinnis), in front of a gorgonian sea fan (Gorgonia sp)Scalefin anthias / Scalefin basslet (Pseudanthias squamipinnis), in front of a gorgonian sea fan (Gorgonia sp.), Fiji, Pacific Ocean
Pair of Eurasian bee-eaters (Merops apiaster) perched on branch, Pusztaszer reserve, Hungary. May
Male Swallowtail butterfly (Papilio machaon) eating from Wood cranesbill (Geranium sylvaticum), Finland. June
Yellowline arrow crab (Stenorhynchus seticornis) living in association with Giant Caribbean sea anemone (Condylactis gigantea), Dominica, Caribbean
Bobtail squid (Heteroteuthis hawaiiensis) on the seabed, Komodo National Park, Indonesia, Pacific Ocean
Female Goldenrod crab spider (Misumena vatia) camouflaged on Yellow flag iris (Iris pseudacorus) flower waiting for prey to land, Kenfig National Nature Reserve, Glamorgan, Wales, UK, June
RF - Eschmeyers scorpionfish (Rhinopias eschmeyeri) composite image showing different colour variations on black background. Lembeh Strait, North Sulawesi, Indonesia
Mimic leatherjacket / Blacksaddle mimic filefish (Paraluteres prionurus) adult and juvenile, composite image on black background, Bali, Indonesia, Indo-Pacific
Honeycomb / Cushion starfish (Pentaceraster alveolatus) composite image on black background showing colour variations Malapascua Island, Philippines, Indo-Pacific species
Mandarin duck (Aix galericulata) drake swimming on pond, backlit by rising sun, Richmond Park, London, UK. February
Close up of section of Swallowtail butterfly (Papilio machaon) caterpillar feet gripping stem of Wild carrot (Daucus carota). The Netherlands. August
Three Grey partridge (Perdix perdix) walking in snow. Near Nijmegen, the Netherlands. February
Group of Grey partridge (Perdix perdix) huddled for warmth in snowy field. Near Nijmegen, the Netherlands. February
Ventral view of Swallowtail butterfly (Papilio machaon) in flight. The Netherlands. July
Swallowtail butterfly (Papilio machaon) nectaring on thistle (Cirsium sp. ) flower, with cyclists in backgroundSwallowtail butterfly (Papilio machaon) nectaring on thistle (Cirsium sp.) flower, with cyclists in background. The Netherlands. July
RF - Two Swallowtail butterflies (Papilio machaon) flying to feed on thistle (Cirsium sp. ) bush in flowerRF - Two Swallowtail butterflies (Papilio machaon) flying to feed on thistle (Cirsium sp.) bush in flower. The Netherlands. July
Mandarinfish (Synchiropus splendidus) leaving the protection of finger coral to look for a mate at dusk, Yap, Micronesia, Pacific Ocean
Slate pencil sea urchin (Heterocentrotus mammillatus) on a reef with Black triggerfish (Melichthys niger) and a Bluestripe snapper (Lutjanus kasmira) swimming above, Maui, Hawaii, Pacific Ocean
Detail of octo coral polyps from Sea fan (Gorgonia sp. ), Triton Bay, West Papua, Indonesia, Pacific OceanDetail of octo coral polyps from Sea fan (Gorgonia sp.), Triton Bay, West Papua, Indonesia, Pacific Ocean
Cleaning shrimp (Periclimenes sp) on a coral reef, close up, Triton Bay, West Papua, Indonesia, Pacific Ocean
Soft coral (Dendronepthya sp) garden, Iris Strait, Triton Bay, Indonesia, Pacific Ocean
Pair of Mandarinfish (Synchiropus splendidus) mating, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Apennine treacle mustard (Erysimum pseudorhaeticum) and Greater milkwort (Polygala major) in flower in dry field, Gran Sasso, Abruzzo, Italy. June
Fallow fields burst into flower containing Corn chamomile (Anthemis arvensis), Hoarry mullein (Verbascum pulverulentum), Purple toadflax (Linaria purpurea) and Common poppy (Papaver rhoeas), Abruzzo
Fallow field in flower containing Poppy (Papaver rhoeas), Woad (Isatis tinctoria), Corn chamomile (Anthemis arvensis) Olympian mullein (Verbascum longifolium) and Purple toadflax (Linaria purpurea)
