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Cotton or hibiscus harlequin bug (Tectocoris diophthalmus) nymph feeding on a green leaf, Brisbane, Queensland, Australia. April
Selection of wildflowers against white background, including Thistle (Cirsium), Common rockrose (Helianthemum chamaecistus)
Common octopus (Octopus vulgaris) on a coral reef, The Bahamas. August
Annual Honesty (Lunaria annua) seed heads arranged in white background
Hyacinth Macaw (Anodorhynchus hyacinthinus) in forest bordering of the Cuiaba River, Northern Pantanal, Brazil. September
Congo cockatoo (Impatiens niamniamensis) flower, backlit with nectar level visible in spur. Cultivated in glasshouse. Native to Africa
Aerial view of autumn mixed forest tree canopy, Laurentides forest, Quebec, Canada, September 2001
Fly Agaric Fungus (Amanita muscaria) within forestry plantation, Strathspey, Scotland, UK, September
Rainbow lorikeet (Trichoglossus moluccanus) on a branch, Cania Gorge National Park, Queensland, Australia. September
Eastern bluebird (Sialia sialis) male fluttering wings while perched on cherry (Prunus sp.) branch in spring, New York, USA, April
Rufous motmot (Baryphthengus martii) perched on branch. Rancho Naturalista, Cartago, Costa Rica
Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica
Red-legged honeycreeper (Cyanerpes cyaneus) two males on barbed wire. Costa Rica
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
Male Golden Pheasant (Chrysolophus pictus) standing on grass in display plumage. Kew Gardens, England, UK. March
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Close-up up of lips of giant clam (Tridacna gigas), Palau, Micronesia
Sheet ice crystals viewed by polarised light
Fluorescent coral photographed with ultraviolet / UV light, Bonaire, Leeward Antilles, Caribbean region, Netherlands Antilles
Detail of Ocellated Turkey (Meleagris / Agriocharis ocellata) plumage. El Mirador-Rio Azul National Park, Department of Peten, Guatemala
Bullseye / Io moth (Automeris io) showing eye spot markings on wings during deimatic display to deter predators, photographed on a white background, originating from North and Central America
Indian moon / Indian luna moth (Actias selene) head-on view showing feather-like antennae. Captive
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Red-and-green macaw (Ara chloropterus) flock feeding on wall of clay lick. Manu Biosphere Reserve, Amazonia, Peru
Kaleidoscopic image of soft corals (Dendronephthya sp). Egypt, Red Sea
Stove-pipe sponge (Aplysina archeri) and Orange elephant ear sponge (Agelas clathrodes) Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean
Magnificent sea anemone (Heteractis magnifica) with Anemone fish within tentacles, Indonesia. Indo-Pacific
Amphipod (Iphimedia obesa) on top of a colony of Deadmans fingers (Alcyonium digitatum) coral, Loch Carron, Ross and Cromarty, Scotland, UK, April
UK Ladybird species, native and invasive, from left to right: Seven-Spot (Coccinella 7-punctata), Harlequin (Harmonia axyridis), Harlequin morph, Two-Spot (Adalia bipunctata)
Small elephant hawk-moth (Deilephila porcellus) on stem. Captive, larvae collected previous year. Downings, County Donegal, Republic of Ireland. May
Red admiral butterfly (Vanessa atalanta) on Buddleia flowers, County Down, Northern Ireland, UK, August
Red-eyed tree frog {Agalychnis callidryas} resting in Bromeliad flower, Nicaragua, June
Fiery throated hummingbird (Panterpe insignis) an endemic bird species. Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
The Tokay gecko (Gekko gecko) licking its eye, captive, from Asia
Rainbow lorikeet (Trichoglossus haematodus) Queensland, Australia
Cairns birdwing butterfly (Ornithoptera priamus) male on fern leaf and Blue Mountain Swallowtail butterfly in flight, Atherton Tablelands, Queensland, Australia, October
Blue jay (Cyanocitta cristata) perched amongst autumn foliage, New York, USA, October
Male Black faced blenny (Tripterygion delaisi), part of a sequence showing colour change, Swanage, Dorset, England, UK, May. 2/4
John dory (Zeus faber), Porthkerris Cove, Cornwall, England, UK, June
Tompot blenny (Parablennius gattorugine) beneath Swanage Pier, Dorset, UK. Photographed during May. English Channel
Old fishing boat held upright at low tide with ropes and chains, Roa Island, Morecambe Bay, Cumbria, England, UK, February 2012
Wood warbler (Phylloscopus sibilatrix) in flight in sesile oak forest, Wales, UK, May. 2020VISION Book Plate
Adult Atlantic puffin (Fratercula arctica) in flight in summer, backlit, Isle of Lunga, Inner Hebrides, Scotland, UK, July 2011. 2020VISION Book Plate
The Cuckmere River at sunset, Seven Sisters Country Park, South Downs National Park, East Sussex, England, UK, July 2011
Goldfinch (Carduelis carduelis) adult feeding on teasel seeds, UK, February
Lilac-breasted roller (Coracias caudata) stretching wings on acacia branch, Masai Mara Game Reserve, Kenya. November
A misty morning over a mixed woodland in autumn, Kinnoull Hill Woodland Park, Perthshire, Scotland, November 2011. Highly commended in the Wild Woods Category of the British Wildlife Photography