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Eyelash viper (Bothriechis schlegelii) waiting on Heliconia flower (Heliconia lankasteri) to ambush hummingbird prey, Golden colour morph La Selva, Costa Rica
Rainbow eucalyptus (Eucalyptus deglupta) bark, Costa Rica
Strawberry poison dart frog (Oophaga / Dendrobates pumilio) in cup fungus, La Selva Field Station, Costa Rica
Honeybee (Apis mellifera) flying to Bluebell flowers (Hyacinthoides non-scripta) Monmouthshire, Wales, UK. April
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
European honey bee (Apis mellifera) in flight, feeding on flowers (Brachyglottis sp). Monmouthshire, Wales, UK, June
Black veined white butterfly (Aporia crataegi) on grass, Gavarnie-Gedre, Pyrenees National Park, France, June
Geranium Argus butterfly (Plebejus eumedon) feeding on Wood Cranesbill (Geranium sylvaticum) northern Finland, June
Male Ostrich (Struthio camelus) protecting chicks from the sun with its wings, Samburu National Reserve, Kenya, Africa
Grey heron (Ardea cinerea) adult in breeding plumage, close-up of head and colourful orange beak, backlit, Berlin Zoological Garden, Berlin Zoo, Germany. February
Lesser flamingo (Phoeniconaias minor) portrait, Lake Bogoria, Rift valley, Kenya, Africa
Portrait of brown and white coated Holland lop-eared domestic rabbit (Oryctolagus cuniculus domesticus), Lorraine, France, July
Nosy Be Pygmy Leaf Chameleon (Brookesia minima) female, worlds smallest reptile. Nosy Be, Madagascar. Photograph taken on location for BBC Wild Madagascar Series, January 2010
Indri (Indri indri) calling, Andasibe-Mantadia NP, Madagascar
Fresh Wych elm (Ulmus glabra) leaves in spring. Dorset, UK May
Fringed leaf frog (Cruziohyla craspedopus) on branch, Yasuni National Park, Orellana, Ecuador
Las gralarias glass frog (Nymphargus lasgralarias) on leaf, Mindo, Pichincha, Ecuador
Shieldhead gecko (Gonatodes caudiscutatus) on branch, Canande, Esmeraldas, Ecuador
Marine Iguana (Amblyrhynchus cristatus) perched on a rock. Floreana Island, Galapagos Islands, Ecuador
Juvenile of Amazon Tree-Boa (Corallus hortulanus) in defensive position. Yasuni National Park, Orellana, Ecuador
Metallic leaf beetle ( Eumolpinae) with rain droplets, in Aiuruoca, Minas Gerais, Brazil. South-east Atlantic forest
Scales of a Owl butterfly (Caligo sp) magnified 28x, deceased insect found in Tapirai Sao Paulo, Brazil. South-east Atlantic forest
Close-up of a Metallic jewel beetle (Buspretidae) in Aiuruoca, Minas Gerais, Brazil. South-east Atlantic forest
Metallic leaf beetle (Chrysomelidae) with rain droplets, frontal view, in Aiuruoca, Minas Gerais, Brazil. South-east Atlantic forest
Raja Brookes Birdwing Butterfly (Trogonoptera brookiana), Borneo
Rangimairewhenua or Blue Lake, Nelson Lakes National Park, Southern Alps, New Zealand, February 2013. Photographed for The Freshwater Project
Laguna Miluni, contaminated with mine effluent and with a low level due to drought. This reservoir is fed by glacial meltwater from the Andean peak, Huayna Potosi, Bolivian Andes. October 2015
RF -Three colourful Red-and-green macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
RF - Four colourful Red-and-green macaws or Green-winged macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
Wavy-lined emerald moth caterpillar (Synchlora aerata) which camouflages itself by attaching parts of flower to itself, on Black-eyed susan (Rudbeckia hirta) Morris Arboretum, Philadelphia
Dogbane beetle (Chrysochus auratus) on dogbane, Crossways Preserve, Philadelphia, Pennsylvania, USA, August
Passionflower (Passiflora cubalensis) Yanacocha Reserve, Ecuador
Red-and-green macaw (Ara chloropterus) in flight over forest canopy. Buraco das Araras, Jardim, Mato Grosso do Sul, Brazil. September
Snowy owl (Bubo scandiaca) female face close up, Canada February
Great Grey Owl (Strix nebulosa) landing in snow, Kuusamo, Finland
RF- Indian eagle owl (Bubo bengalensis) head portrait, captive, from India
RF- Indian eagle owl (Bubo bengalensis) portrait, captive, from India
Steppe eagle (Aquila nipalensis) in flight, Oman, November
RF- Golden eagle (Aquila chrysaetos) landing in snow, Flatanger, Norway. November
Meadow of Fragrant Orchids (Gymnadenia conopsea) at sunset. Tirol, Austrian Alps, Austria, June
European / Common Lobster (Homarus gammarus) in rocky retreat. Controlled conditions, Cornwall, UK. May
Goldenrod crab spider (Misumena vatia) camouflaged on yellow flower. Nordtirol, Austrian Alps, July
Ash Tree (Fraxinus excelsior) growing in a field. Peak District National Park, Derbyshire, UK. August
RF - Red Deer stags (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Swallowtail butterfly (Papilio machaon) in garden, Norfolk, England, UK, June
RF - Female emperor dragonfly (Anax imperator) laying eggs on garden pond, Broxwater, Cornwall, UK. July
Emperor moth (Saturnia pavonia) caterpillar feeding on Ling heather (Calluna vulgaris) taken with fish eye view, Peak District National Park, Derbyshire, UK. August
Great crested newt (Triturus cristatus) male in breeding condition, amongst Soft hornwort (Ceratophyllum submersum), Herefordshire, England, UK, May. Captive