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Bichon frise beside Colorado blue columbine (Aquilegia coerulea) flowers. Old Lyme, Connecticut, USA. May
White witch moth (Thysania agrippina) camouflaged against bark of tree. Manu Biosphere Reserve, Amazonia, Peru
Bichon frise standing on rock amongst Rhododendron flowers. Haddam, Connecticut, USA. May
Snowy egret (Egretta thula) portrait, viewed from below. St. Petersburg, Florida, USA
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Red-and-green macaw (Ara chloropterus) flock feeding on wall of clay lick. Manu Biosphere Reserve, Amazonia, Peru
RF - Maidens veil / Bridal veil fungus (Phallus indusiatus) with indusium fully formed, on rainforest floor. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Wild boar (Sus scrofa) in old mixed conifer and broadleaf forest, Punia Forest Reserve, Lithuania, May
Aurora Borealis over Vestrahorn, Stokksnes, Hofn, Iceland. March 2018
Vik i Myrdal church in winter, with full moon, Iceland. January 2015
Kaleidoscopic image of soft corals (Dendronephthya sp). Egypt, Red Sea
Stove-pipe sponge (Aplysina archeri) and Orange elephant ear sponge (Agelas clathrodes) Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean
RF - Red Dachshund puppy lying down
RF - Blue tabby kitten, 12 weeks, jumping up
Silver tabby kitten, Freya, age 8 weeks
Silver tabby Persian-cross kitten arching back in playful confrontation
Black-and-tan Dachshund puppy walking
Northern raven (Corvus corax) pair perching on branch. Danube Delta, Romania, May
Great white egret (Ardea alba) group of three in winter, Kiskunsag National Park, Hungary. Winner of the Bird Category of Nature Photographer of the Year Awards 2018
Young White rhinoceros (Ceratotherium simum) in a reinforced steel boma, blindfolded and partially drugged after a long journey from South Africa, before being released into the wild in Botswana
Blue butterfly (Lycaenidae) frozen in ice, Cortes de la Frontera, Los Alcornocales Natural Park. Spain
Aerial view of Amazon Rainforest and the Yavari River, and oxbow lake, Peru. July 2015
UK Ladybird species, native and invasive, from left to right: Seven-Spot (Coccinella 7-punctata), Harlequin (Harmonia axyridis), Harlequin morph, Two-Spot (Adalia bipunctata)
Chinese moon moth (Actias dubernardi, female at rest amongst leaves. Dayaoshan, Jinxin, Guangxi, China
Small elephant hawk-moth (Deilephila porcellus) on stem. Captive, larvae collected previous year. Downings, County Donegal, Republic of Ireland. May
Scarlet elf cup fungus (Sarcoscypha coccinea) amongst Opposite-leaved golden-saxifrage (Chrysosplenium oppositifolium). Clare Glen, Tandragee, County Armagh. March
Shag (Phalacrocorax aristotelis) in flight over sea. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Marsh fritillary (Euphydryas aurinia) on fern. South of Casteil, Pyrenees Orientales, south west France. May
Mountain clouded butterfly (Colias phicomone) on clover flower, Stelvo Pass, Alps, Italy, June
Saturniid moth (Eacles ormondei), female camouflaged in leaf litter. Izabal, Guatemala
Oleander Hawkmoth (Daphnis nerii) resting, Southern Sicily, Italy. March
Ghost glass frog (Sachatamia ilex), Costa Rica
Snowy owl (Bubo scandiacus) in flight, Ontario, Canada, January
Thread-tail dartgoby (Ptereleotris hanae) Sharp Island or Kiu Tsui Chau, Kiu Tsui Country Park, Hong Kong, China
Paddle-flap scorpionfish (Rhinopias eschmeyeri) Puerto Galera, Philippines
RF - White lion (Panthera leo) male, portrait of head. Captive, Netherlands
Leaves of a Ginkgo tree or Maidenhair tree (Ginkgo biloba) Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Polar bear (Ursus maritimus) juvenile crossing over newly forming pack ice, during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
RF- Maine Coon mother cat, with two kittens, 7 weeks
Northern lights over Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
RF - Giant panda (Ailuropoda melanoleuca) in snow, captive
RF - Polar Bears (Ursus maritimus) males fighting, Churchill, Canada, November
Grey Arabian stallion running in desert dunes near Dubai, United Arab Emirates
RF - Head and mane of grey Andalusian stallion running in bullfighting arena, Southern Spain, Europe
Two wild pinto Mustang stallions battle for dominance in Sand Wash Basin, Colorado, USA
Young Cremello Lusitano stallion running through field of wildflowers in southern Spain, Europe. April
Two white Camargue horses in ocean of Camargue, France, Europe. May
RF - Head portrait of grey Andalusian mare with long mane in snow, Berthoud, Colorado, USA. January