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Oxlips (Primula eliator) flowering in coppice woodland, a rare and important ancient woodland indicator species, Suffolk, England, UK, April
Green Veined White Butterfly (Pieris napi) Roosting on Bluebell flower, Cambridgeshire, England, UK, April
Burrowing owl (Athene cunicularia) doing rain dance, spreading wings and dancing while it rained, Arizona, USA
Blackcap (Sylviya atricapilla) male perched in blossom, Hungary, April
Arctic fox (Vulpes lagopus), juvenile licking lips, winter pelage. Dovrefjell National Park, Norway. September
Burrowing owl (Athene cunicularia) rotating head, Arizona, USA
Australian green tree frog (Litoria caerulea) camouflaged on Palm leaf. Lake Argyle, Kununurra, Western Australia
Melanistic leopard / Black panther (Panthera pardus) Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India
Great grey owl (Strix nebulosa) taking off from snowy ground, Kuusamo, Finland. March
Red-legged honeycreeper (Cyanerpes cyaneus) two males on barbed wire. Costa Rica
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
Male Golden Pheasant (Chrysolophus pictus) standing on grass in display plumage. Kew Gardens, England, UK. March
Flowers (Trollius sp) Balang Mountain, Wolong National Nature Reserve, Sichuan Giant Panda Sanctuaries - Wolong, Mt Siguniang and Jiajin Mountains UNESCO World Heritage Site, Sichuan Province, China
Black grouse (Lyrurus tetrix), male at lek, Finland, April
Saffron crocuses (Crocus sativus), cultivated for saffron, Lleida, Catalonia, Spain, November
Soft corals (Dendronephthya sp) in coral reef, West Papua, New Guinea
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Canebreak treefrog (Aplastodiscus sp) Guainumbi Private Reserve, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil
Scarlet ibis (Eudocimus ruber), two birds perched on bushes, Coro, Venezuela. Small repro only
Scarlet ibis (Eudocimus ruber), in flight, Coro, Venezuela
Nudibranchs (Nembrotha chamberlaini) feeding on Tunicates (sea squirt) on a coral reef. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Southern Viscacha (Lagidium viscacia) at rest, Isla Incahuasi in the Salar de Uyuni, altiplano, Bolivia
Emerald toucanet (Aulacorhynchus prasinus) Rio Platano Biosphere Reserve and UNESCO World Heritage Site, La Mosquitia, Honduras
Deep sea jellyfish (Periphylla periphylla), Trondheimsfjord, Norway, July
Black smoke fluffy Lionhead x Lop rabbit
Patterdale terrier dog puppy, Korka, age 4 months
Strawberry poison frog (Oophaga pumilio) photographed on a white background in mobile field studio, Central Caribbean foothills, Costa Rica
Sheet ice crystals viewed by polarised light
Fluorescent coral photographed with ultraviolet / UV light, Bonaire, Leeward Antilles, Caribbean region, Netherlands Antilles
Aerial view of shifting sand banks off the coast of the Bazaruto Archipelago, Mozambique, May 2011
Abstract photograph of grasses and yellow flowers, Vosges, France, June
Dalmatian pelicans (Pelecanus crispus) underwater view of group feeding, Lake Kirkini, Greece. Highly commended in the GDT competition 2013
Ice formations, Lake Sassolo, Sambuco valley, Ticino, Switzerland
RF- Aerial view of the Okavango delta, Botswana, May 2010
Jersey tiger moth (Euplagia quadripunctaria) with less common yellow colour variation feeding on ragwort, Lewisham, London, UK, August
Red beetle (Pyrochroidae) sitting on a leaf, Tongbiguan Nature Reserve, Dehong prefecture, Yunnan province, China, May
Moss covered fallen trees in old mixed conifer and broadleaf forest, Punia Forest Reserve, Lithuania, May
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
Great barracuda (Sphyraena barracuda) with three parasitic Copepods at end of upper jaw. Hawaii
Oreocallis grandiflora (Proteaceae family), Manu Cloud forest at 3500 metres altitude, Peru. November
Scots pine (Pinus sylvestris) forest with Bracken (Pteridium aquilinum) ground flora in autumn, Glenfeshie, Cairngorms National Park, Scotland, UK, October
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
Seedling cone plants (Conophytum subfenestratum), approximately two months after sowing. Focus-stacked image
Chesapeake Bay retriever male standing in saltmarsh, Long Island Sound, Madison, Connecticut, USA
English springer spaniel in Rhododendron. Haddam, Middlesex, Connecticut, USA
Parson's chameleon (Calumma parsonii), male hunting in forest canopy. Mitsinjo Forest, Andasibe-Mantadia National Park, eastern Madagascar
Green-veined white butterfly (Pieris napi) Whitelye Common Nature Reserve, Monmouthshire, Wales, UK