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Grey heron (Ardea cinerea) in rain with another heron behind, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Two Burrowing owls (Athene cunicularia) fledgling with adult about to take off, Pantanal, Brazil
Pleiades or Seven Sisters (Messier 45 aka M45) in Taurus Constellation, taken from Eastern Colorado, USA. 8 November 2015
Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April
Leaf cutter ants (Atta sp) carrying colourful plant matter, reflected in water, Laguna del Lagarto, Santa Rita, Costa Rica
Red fox (Vulpes vulpes) vixen in front of Clifton Suspension Bridge at night. Avon Gorge, Bristol, UK. March 2016
Kottigehara dancing frog (Micrixalus kottigeharensis), dancing frog name given because of foot flagging behavior between competing males. Endemic to Western Ghats. Agumbe, Karnataka, India
Robin (Erithacus rubecula) resting on spruce sapling, Black Isle, Scotland, UK April
Fiordland crested penguins (Eudyptes pachyrhynchus) in dense coastal forest, Lake Moeraki, South Island, New Zealand, November, threatened species
Red fox (Vulpes vulpes) at night in snow, camera trap image, Jura Mountains, Switzerland, August. Commended in the Mammals category of the Asferico Competition 2016
Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
North American Bison (Bison bison) coated in frost standing on snow, Yellowstone National Park, Wyoming, USA, January
Tree ferns (Cyatheaceae) in Kells Bay Gardens, Ring of Kerry, Iveragh Peninsula, County Kerry, Ireland, Europe. September 2015
Murmuration of Common starlings (Sturnus vulgaris) with Peregrine falcon (Falco peregrinus) hunting them, The Netherlands, November
Whooper swans (Cygnus cygnus) three coming into land, Hokkaido, Japan, February
Moose (Alces alces) bull swimming in water, Baxter State Park, Maine, USA
Angular roughshark (Oxynotus centrina) a deepsea species living at 80-300m depth, caught by fishermen and released, Costa Brava, Catalunya, Spain, Mediterranean Sea
Marsh fritillary butterfly (Euphydryas aurinia) feeding on Wild garlic / Ramsons (Allium ursinum) at Strawberry Banks, Gloucestershire Wildlife Trust (GWT), Nature Reserve, Gloucestershire, UK. May
Coconut / Veined octopus (Amphioctopus marginatus) hunts in the sand at night, while illuminated by a divers torch beam, Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Pacific Ocean
Bearded tit (Panurus biarmicus) male portrait feeding, Helsinki, Finland, January
Tabby kitten, 6 weeks, lying with head up and raised paw
Young girl in traditional dress. Mursi tribe, Mago National Park. Omo Valley, Ethiopia
Sunset over Loch Coire Mhic Fhearchair from Beinn Eighe with the sky reflected in the water. Torridon, Highlands, Scotland, UK, June 2016
Bison (Bison bison) herd walking in snow, Yellowstone National Park, Wyoming, USA, February
Blue whale (Balaenoptera musculus) underwater, Coronado Islands, Baja California, Mexico, Eastern Pacific Ocean
Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
Common loon (Gavia immer) nesting at edge of lake and calling, Acadia National Park, Maine, USA, August
Great egret (Ardea alba) two fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Wild Eurasian lynx (Lynx lynx) at night with city lights and sky glow behind, Switzerland. December. Taken with remote camera trap
African wild dog (Lycaon pictus) interacting with pup age two months, Okavango Delta, Botswana
Alpine ibex (Capra ibex), male in habitat, Bernese Alps, Switzerland. July
Zanzibar red colobus monkey (Procolobus kirkii) crossing the main road through the Reserve, Jozani forest, Jozani Chwaka Bay NP, Zanzibar, Tanzania, August
African Wild dogs or Cape hunting dogs (Lycaon pictus) at close range taken from ground level, Zimanga Private Game Reserve, South Africa
Cheetah (Acinonyx jubatus) staring back over its shoulder, black and white. Save Valley Conservancy, Zimbabwe, August
Juvenile White tailed eagle (Haliaeetus albicilla)in flight, with Hooded crow (Corvus cornix) mobbing it, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Aggregation of Golden jellyfish (Mastigias sp.) in a marine lake in Palau, the golden colour of this species comes from symbiotic algae in its tissues
Rainforest and valley in the Sarawaget range, Huon Peninsula, Papua New Guinea. November 2006
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Lava lake at night in the crater of Nyiragongo Volcano, Virunga National Park, North Kivu Province, Democratic Republic of Congo
Patterdale terrier dog puppy, Korka, age 4 months
Cow killer velvet ant (Dasymutilla occidentalis) on white background, Tuscaloosa County, Alabama, USA September
Sperm whale (Physeter macrocephalus) female at the ocean surface with her mouth open. This whale has just surfaced from foraging in deep water. Ogasawara or Bonin Islands, Japan
Red bat (Lasiurus borealis) female flying; shots taken with high speed flash San Saba County, Texas, USA. Controlled conditions. July
Blackbird (Turdus merula) in flight to feed on berries, Helsinki, Finland. November
Composite photograph showing diversity of colour and pattern in samples of granite
Green iguana (Iguana iguana) rear view of basking in sun, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico, May
American crocodile (Crocodylus acutus) rear view of animal resting in shallow water with head at surface, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico
Ptarmigan (Lagopus mutus) in snowy landscape, Utsjoki, Finland, February
Bearded reedling (Panurus biarmicus) on frosty reed seed head, Helsinki, Finland, January
Part of a submerged Holocene forest, revealed after a winter storm from beneath the sand at Hells Mouth (Porth Neigwl), Abersoch Wales, UK March 2014
Great grey owl (Strix nebulosa) taking off from snowy ground, Kuusamo, Finland. March
Thresher shark (Alopias pelagicus) swimming over seabed to be cleaned by cleaner wrasses, Cebu, Malaspascua, Philippines, September
Red fox (Vulpes vulpes) resting with eyes open, Vanoise National Park, Rhone Alpes, France. October
Shore of lake in Torres del Paine. Patagonia, Puerto Natales, Chile
Blacksmiths (Chromis punctipinnis) school swam through a giant kelp (Macrocystis pyrifera) forest, Santa Barbara Island, Channel Islands. Los Angeles, California, USA, North East Pacific Ocean
Grey seal (Haichaoerus grypus) swimming below surface with sunbeams. Lundy Island, Devon, UK. Bristol Channel, August
Capercaillie (Tetrao urogallus) male displaying in rain, Kuusamo, Finland. March
Red-legged honeycreeper (Cyanerpes cyaneus) two males on barbed wire. Costa Rica
Flock of Tree sparrows (Passer montanus) in flight, with a Greenfinch (Chloris chloris) blurred motion, Pacsmag, Hungary
Bauxite mine, blue water caused by industrial runoff, minerals into local water, Linden town, Guyana, South America
Lava flow from Kilauea Volcano flowing into the Pacific Ocean, Kalapana Coast, Big Island, Hawaii. August 2016
A Cotswold Lion sheep, rare breed, Gloucestershire, UK. June
Aerial view of Belas Knap, a neolithic chambered long barrow on the Cotswold Way, Winchcombe, Gloucestershire, UK. Shot by aerial drone with Ca permit. January 2016
Nurse shark (Ginglymostoma cirratum) and Horse-eye jacks (Caranx latus) Shark Ray Alley, Hol Chan Marine Reserve, Belize Barrier Reef, Belize
Whale shark (Rhincodon typus) viewed from above, Cenderawasih Bay, West Papua. Indonesia
Skeleton of a seahorse (Hippocampus sp. ) showing the remarkable structure of bonesSkeleton of a seahorse (Hippocampus sp.) showing the remarkable structure of bones that form a virtual exoskeleton, with few deep internal bones, in this unusual vertebrate group
Historical illustration of Common octopus (Octopus vulgaris) Pierre Denys de Montfort engraving from Histoire Naturelle Generale et Particuliere des Mollusques, 1801-1802
Aerial view of Giant water lily (Victoria amazonica) leaves in river, i Rurununi savanna, Guyana, South America
Mount Roraima tepui (flat top mountain) is the highest of the Pakaraima chain of Tepuis plateaus in South America, Pakarajma Mountains, near Phillipai, West Guyana, South America
Mexican free-tailed bats, (Tadarida brasiliensis), leaving Bracken Cave, Texas
Galapagos racer snakes (Pseudalsophis biserialis) group alert watching for prey on beach, Galapagos, June. Endangered species. Taken on location for BBC tv Blue Planet 2 series
Sable antelope (Hippotragus niger) bull charging through the shallows of the Selinda Spillway, with another behind, northern Botswana, August
Storm waves off the coast of Praia do Norte, Nazare, Portugal. January
Flock of Mallard ducks (Anas platyrhynchos) taking off, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. December
European robin (Erithacus rubecula) low angle view near church, Bristol, UK. January
Low angle view of Porcelain fungus (Oudemansiella mucida) growing on a dead Beech tree (Fagus sylvatica). Plitvice Lakes National Park, Croatia. November
Black faced sheep ram with twisted horns, Mull, Scotland, UK. January
Zebra (Equus quagga) leaping during stampede, Serengeti, Tanzania. Vignette added and right edge expanded
European hamster (Cricetus cricetus) male, in underground burrow, captive
Puma (Puma concolor) in high altitude habitat of Torres del Paine National Park, Chile
Night-blooming cereus cactus (Acanthocereus tetragonus), flower bud opening at night, Texas, USA. Sequence 7 of 7. August
Malabar pit viper (Trimeresurus malabaricus), green colour morph. Agumbe, Karnataka, Western Ghats, India. Endemic
Malabar torrent toad (Ghatophryne ornata) with bright yellow-orange spots more on abdomen. Coorg, Karnataka, India. Endemic to the Western Ghats. Endangered species. Endangered species
Giant Manta (Manta birostris) with Remoras (Remora brachyptera) attached to both wings, offshore Santa Maria Island, Azores, Atlantic Ocean