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Leaf cutter ants (Atta sp) carrying colourful plant matter, reflected in water, Laguna del Lagarto, Santa Rita, Costa Rica
Red fox (Vulpes vulpes) vixen in front of Clifton Suspension Bridge at night. Avon Gorge, Bristol, UK. March 2016
Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April
Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
Angular roughshark (Oxynotus centrina) a deepsea species living at 80-300m depth, caught by fishermen and released, Costa Brava, Catalunya, Spain, Mediterranean Sea
Red fox (Vulpes vulpes) at night in snow, camera trap image, Jura Mountains, Switzerland, August. Commended in the Mammals category of the Asferico Competition 2016
Pleiades or Seven Sisters (Messier 45 aka M45) in Taurus Constellation, taken from Eastern Colorado, USA. 8 November 2015
Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
Tree ferns (Cyatheaceae) in Kells Bay Gardens, Ring of Kerry, Iveragh Peninsula, County Kerry, Ireland, Europe. September 2015
Murmuration of Common starlings (Sturnus vulgaris) with Peregrine falcon (Falco peregrinus) hunting them, The Netherlands, November
Whooper swans (Cygnus cygnus) three coming into land, Hokkaido, Japan, February
Young girl in traditional dress. Mursi tribe, Mago National Park. Omo Valley, Ethiopia
Moose (Alces alces) bull swimming in water, Baxter State Park, Maine, USA
Fiordland crested penguins (Eudyptes pachyrhynchus) in dense coastal forest, Lake Moeraki, South Island, New Zealand, November, threatened species
Great egret (Ardea alba) two fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
African wild dog (Lycaon pictus) interacting with pup age two months, Okavango Delta, Botswana
Alpine ibex (Capra ibex), male in habitat, Bernese Alps, Switzerland. July
African Wild dogs or Cape hunting dogs (Lycaon pictus) at close range taken from ground level, Zimanga Private Game Reserve, South Africa
Bearded tit (Panurus biarmicus) male portrait feeding, Helsinki, Finland, January
Tabby kitten, 6 weeks, lying with head up and raised paw
Cheetah (Acinonyx jubatus) staring back over its shoulder, black and white. Save Valley Conservancy, Zimbabwe, August
Sunset over Loch Coire Mhic Fhearchair from Beinn Eighe with the sky reflected in the water. Torridon, Highlands, Scotland, UK, June 2016
Bison (Bison bison) herd walking in snow, Yellowstone National Park, Wyoming, USA, February
Blue whale (Balaenoptera musculus) underwater, Coronado Islands, Baja California, Mexico, Eastern Pacific Ocean
Wild Eurasian lynx (Lynx lynx) at night with city lights and sky glow behind, Switzerland. December. Taken with remote camera trap
Coconut / Veined octopus (Amphioctopus marginatus) hunts in the sand at night, while illuminated by a divers torch beam, Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Pacific Ocean
Zanzibar red colobus monkey (Procolobus kirkii) crossing the main road through the Reserve, Jozani forest, Jozani Chwaka Bay NP, Zanzibar, Tanzania, August
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Common loon (Gavia immer) nesting at edge of lake and calling, Acadia National Park, Maine, USA, August
Aggregation of Golden jellyfish (Mastigias sp.) in a marine lake in Palau, the golden colour of this species comes from symbiotic algae in its tissues
Lava lake at night in the crater of Nyiragongo Volcano, Virunga National Park, North Kivu Province, Democratic Republic of Congo
Cow killer velvet ant (Dasymutilla occidentalis) on white background, Tuscaloosa County, Alabama, USA September
Black-and-white kitten, Solo, 7 weeks, lying on his side
Sperm whale (Physeter macrocephalus) female at the ocean surface with her mouth open. This whale has just surfaced from foraging in deep water. Ogasawara or Bonin Islands, Japan
Red bat (Lasiurus borealis) female flying; shots taken with high speed flash San Saba County, Texas, USA. Controlled conditions. July
Blackbird (Turdus merula) in flight to feed on berries, Helsinki, Finland. November
Composite photograph showing diversity of colour and pattern in samples of granite
Green iguana (Iguana iguana) rear view of basking in sun, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico, May
American crocodile (Crocodylus acutus) rear view of animal resting in shallow water with head at surface, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico
Bearded reedling (Panurus biarmicus) on frosty reed seed head, Helsinki, Finland, January
Great grey owl (Strix nebulosa) taking off from snowy ground, Kuusamo, Finland. March
Thresher shark (Alopias pelagicus) swimming over seabed to be cleaned by cleaner wrasses, Cebu, Malaspascua, Philippines, September
Red fox (Vulpes vulpes) resting with eyes open, Vanoise National Park, Rhone Alpes, France. October
Shore of lake in Torres del Paine. Patagonia, Puerto Natales, Chile
Blacksmiths (Chromis punctipinnis) school swam through a giant kelp (Macrocystis pyrifera) forest, Santa Barbara Island, Channel Islands. Los Angeles, California, USA, North East Pacific Ocean
Grey seal (Haichaoerus grypus) swimming below surface with sunbeams. Lundy Island, Devon, UK. Bristol Channel, August
Capercaillie (Tetrao urogallus) male displaying in rain, Kuusamo, Finland. March
Red-legged honeycreeper (Cyanerpes cyaneus) two males on barbed wire. Costa Rica