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Corncrake (Crex crex) calling at dawn on machair, North Uist, Outer Hebrides, Scotland, UK, June
Red Deer Stag (Cervus elaphus) roaring. Bradgate Park, Leicestershire, UK, October
Red-throated Diver (Gavia stellata) calling, Iceland, June
Red deer (Cervus elaphus) stag bellowing during rut in grassland, doe in background
Bar headed geese (Anser indicus), four in flight, one calling. Coming in to land at breeding site, Qinghai Lake, Qinghai Province, China
Common cranes displaying by woodland {Grus grus} Finland
Red deer (Cervus elaphus) stag roaring during the rut, Leicestershire, UK, October. Non-ex
Red deer (Cervus elaphus) stag bellowing, surrounded by hinds, Surrey, UK, September
Yellow-rumped cacique (Cacicus cela) calling, Madidi NP, Bolivia
Magellanic penguin (Spheniscus magellanicus) calling at the entrance of burrow, courtship display, Carcass Island, Falkland Islands, October
Redshank (Tringa totanus) silhouetted, Iceland, June
Golden eagle (Aquila chrysaetos) male head portrait beak open calling, Kalvtrask, Vasterbotten, Sweden. December
Golden jackal (Canis aureus) howling in grassland. Danube Delta, Romania, May
Belchers / Peruvian band-tailed gull (Larus belcheri) calling, Paracas National Reserve, Peru
Edible frog (Rana esculenta) male calling at pond surface, Klein Schietveld, Brasschaat, Belgium May
Western Scrub-Jay (Aphelocoma californica), adult calling on icy branch of Possum Haw Holly (Ilex decidua) with berries, Hill Country, Texas, USA. February
Snowy owl (Bubo scandiacus) yawning or bill-stretching, Grays Harbor County, Washington, USA December
Prairie Warbler (Dendroica discolor) male singing on branch of tree, with spring leaves, Tompkins County, New York, USA, May
Grey seal (Halichoerus grypus) mating behaviour, Donna Nook, Lincolnshire, UK, November 2008
European green toad (Bufo viridis) calling, vocal sac inflated, with eyes closed, in a pond (at 2, 711m) Adylsu valley, side valley to Baksan valley and Elbrus, Caucasus, Russia, June 2008
European souslik (Spermophilus citellus) calling, Slovakia, Europe, June 2008
Red-crowned Cranes (Grus japonensis) displaying and calling in snow, Hokkaido, Japan, February
Brown Duckling cheeping, against white background
Common / Eurasian crane (Grus grus) calling at sunset, silhouetted. Lake Hornborga, Sweden, April
RF- Grey Wolf (Canis lupus) howling at sunset (digitally enhanced)
Razorbill (Alca torda) pair calling, between soft focus vegetation. Great Saltee, Saltee Islands, Co. Wexford, Ireland, June
Whooper Swans (Cygnus cygnus) on snow, Caerlaverock WWT, Scotland, Solway, UK, January
Andean Gull (Chroicocephalus serranus) calling in flight, Cotopaxi, Ecuador
Great skua (Stercocarius skua) in flight over Gannet (Morus bassanus) colony, looking for small chicks, whilst Gannets call defensively, Shetland Isles, July
Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Couchs Spadefoot (Scaphiopus couchii) adult at night calling with vocal sac inflated. Laredo, Webb County, South Texas, USA, April
Laysan albatross (Phoebastria immutabilis) head portrait calling mate, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, March
Fieldfare (Turdus pilaris) feeding on Rowan berries (Sorbus hupehensis) in winter, Perthshire, Scotland. WINNER: Young Wildlife Photographer of the Year 2010
Arctic tern (Sterna paradisaea) in flight, vocalising, aggresive behaviour, Farne Islands, Northumberland, UK, June
Arctic tern (Sterna paradisaea) in flight, calling, Farne Islands, Northumberland, UK, June
RF - Chinstrap Penguins (Pygoscelis antarcticus) once calling, South Shetland Islands, Antarctica Continent
Lynx (Lynx lynx) cub calling. Hannau, Germany, September. Captive
