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Wrybill (Anarhynchus frontalis) standing shallow water with beak wide open. Lake Ellesmere, South Island, New Zealand. November
RF - Bald eagle (Haliaeetus leucocephalus) calling, captive, occurs in North America
Common starling singing (Sturnus vulgaris) Finland May
Tennessee Warbler (Vermivora peregrina) male perched in a pine tree branch, singing in spring, Alberta, Canada. May
RF- Capercaillie (Tetrao urogallus) male displaying, Inshriach, Cairngorms National Park, Scotland, UK. February
Male Rock ptarmigan (Lagopus mutus) vocalising whilst in flight, showing winter plumage, Utsjoki, Finland, April
Fallow deer (Dama dama) buck bellowing at dawn during the rut, Cheshire, UK, October
Whooper Swan (Cygnus cygnus) on snow, Caerlaverock WWT, Scotland, Solway, UK, January
Crimson-rumped toucanet (Aulacorhynchus haematopygus) calling, Mashpi, Pichincha, Ecuador
Snowy Owl (Bubo scandiaca) with its beak open. Canada, February
RF- Eagle owl (Bubo bubo) calling, raised in captivity and newly released into the wild. Stockholm archipelago, Uppland, Sweden. September
Red deer (Cervus elaphus) stag bellowing. Surrey, UK, October
Robin (Erithacus rubecula) singing, Norfolk, England, UK, February
Chaffinch (Fringilla coelebs) male singing, Norfolk, UK June
Hourglass treefrog (Dendropsophus ebraccatus) male with inflated vocal sac calling at night, Central Caribbean foothills, Costa Rica
Sumatran Orangutan (Pongo abelii) juvenile aged 2.5 years hanging upside down while playing with mother, North Sumatra, Indonesia. Critically Endangered
Red deer (Cervus elaphus) stag bellowing during rut, Derbyshire, UK
Common Raven (Corvus corax) two in flight during snow storm, Kuusamo, Finland, April
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male calling and performing courtship / territorial display in cattail marsh, Mono Lake Basin, California, USA
Red howler monkey (Alouatta seniculus) howling, captive
Black backed jackal {Canis mesomelas} howling, Etosha NP, Namibia
Gentoo penguins (Pygoscelis papua) calling, Cuverville Island. Antarctic Peninsula, Antarctica
Yellow-bellied toad (Bombina variegata) male in pond, calling, Lucerne, Switzerland. July
Little bittern (Ixobrychus minutus) perched in reedbed calling, Valkenhorst nature reserve, Valkenswaard, the Netherlands. May
Bluethroat (Luscinia svecica cyanecula) male, singing as a courtship display, Germany
Ragdoll-cross kitten, aged 3 weeks, meowing, portrait
Galapagos fur seal (Arctocephalus galapagoensis) with mouth open, head portrait, Santiago Island, Galapagos National Park, Galapagos Islands. Endangered
Black Welsh mountain sheep lamb calling. Wales UK, March
Green bee-eater (Merops orientalis) calling, portrait, Bardia National Park, Terai, Nepal
Three Brown noddies / Common noddies (Anous stolidus) trying to steal a fish from a Brown pelican (Pelecanus occidentalis), Santiago Island, Galapagos Islands, Pacific Ocean
Common pheasant (Phasianus colchicus) male flapping wings and calling, Pusztaszer reserve, Hungary. May
Male Western capercaillie (Tetrao urogallus) calling during lek mating, Finland. March
Pair of adult Red-necked grebe (Podiceps grisegena) calling, Finland. May
Jackdaw (Corvus monedula) swooping down calling with claws raised to threaten a Grey squirrel (Sciurus carolinensis) in the entrance to a nest box the bird wants to nest in which is already occupied
Spoonbill (Platalea leucorodia) calling in shallow pond on misty morning, Valkenhorst Nature Reserve, The Netherlands, August
Southern elephant seal (Mirounga leonina) male, rests and yawns on the snow. Right Whale Bay, South Georgia Island
Northern spadefoot (Notaden melanoscaphus) adult male with vocal sac inflated, calling, Howard River sand flats, Northern Territory, Australia, January
Bilingual frog (Crinia bilingua) adult male calling, Adelaide River, Northern Territory, Australia, January
Saddleback anemonefish (Amphiprion polymnus) barks a warning as it guards a clutch of developing eggs (beneath it) laid on a small stone, below a fold in its home
Red deer (Cervus elaphus) in the pine forest bellowing during the rutting season. Barycz Valley Landscape Park, Poland. September
Short-headed treefrog (Dendropsophus brevifrons) calling with inflated vocal sac. Above a rainforest pond at night, Yasuni National Park, Ecuador
Black-bellied bustard (Lissotis melanogaster) male calling at sunrise. Serengeti National Park, Tanzania
African grey hornbill (Lophoceros nasutus) calling in display, perched on tree snag. Janjanbureh, Gambia
RF - Black-browed albatross (Thalassarche melanophrys) calling and in flight. Bay of Isles, South Atlantic. January. (This image may be licensed either as rights managed or royalty free.)
Arctic tern (Sterna paradisaea) adult calling from rock in breeding colony. Isle of May, Scotland, UK, July
Chimpanzee (Pan troglodytes schweinfurthii) male vocalising in tree, Kibale National Park, Uganda, Africa
Waved albatrosses (Phoebastria irrorata) courting, Punta Suarez, Espanola Island, Galapagos
