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Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Flightless cormorant (Phalacrocorax harrisi), adult and two chicks. Beagle Crater, Isabela Island, Galapagos
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Green Woodpecker (Picus viridis ) male, Germany, December
Nightingale (Luscinia megarhynchos ) singing, Germany, April
Common Swift (Apus apus) in flight reflected in water, Norfolk, England, UK. July
Group of Swan geese (Anser cygnoides) on grassland, Inner Mongolia, China
Ptarmigan (Lagopus mutus) in snowy landscape, Utsjoki, Finland, February
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Two rainbow lorikeets (Trichoglossus moluccanus) on branch, Werribee, Victoria, Australia
Little bittern (Ixobrychus minutus) watching Great warbler (Acrocephalus arundinaceus) with prey, Lake Csaj, Kiskunsagi National Park, Hungary
Corncrake (Crex Crex) calling, with breath vapor in early morning, Kiskunsagi National Park, Hungary
Montezuma oropendola (Psarocolius montezuma) hanging upside down as part of a courtship display. Costa Rica
Dalmatian pelican (Pelecanus crispus) close up of feathers, Danube Delta, Romania
Jack snipe (Lymnocryptes minima) feeding in margins of watercress bed, Hertfordshire, England, UK, January
Copper-rumped hummingbird (Saucerottia tobaci) feeding, Tobago, West Indies
White tailed sea eagle (Haliaeetus albicilla) interacting with Great egret (Ardea alba) Lake Csaj, Kiskunsagi National Park, Hungary. January
Great egrets (Ardea alba) sky pointing, sizing each other up before fighting, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Flock of Mallard ducks (Anas platyrhynchos) taking off, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. December
Great egret (Ardea alba) and Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
European robin (Erithacus rubecula) low angle view near church, Bristol, UK. January
Common eider (Somateria mollissima), female on her nest with sea behind, Finland, May
Group of Great egrets (Ardea alba), Grey heron (Ardea cinerea) and Eurasian spoonbill (Platalea leucorodia) at night with stars and moon. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. May
Common raven (Corvus corax) two adults in flight, Hornoya, Norway, April
Adult Kea (Nestor notabilis) close-up portrait. Arthurs Pass National Park, New Zealand, July. Endangered Species
Wood sandpiper (Tringa glareola) in reed bed, Finland, May
Long-tailed tit (Aegithalos caudatus caudatus) placing feather in nest, Finland. April
Blackcap (Sylvia atricapilla) female and two males huddling together for warmth before going to sleep, Finland, May
Ural owl (Strix uralensis), juvenile, Finland, June
Bearded reedling / tit (Panurus biarmicus), male in snow, Finland, March
Lesser spotted woodpecker (Dendrocopos minor), male at nest hole, Finland, June
Caribbean Flamingo (Phoenicopterus ruber) chick age 5 days, walking around nest, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June
Common redpoll (Acanthis flammea), flock feeding, Finland, January
Southern Rockhopper penguin (Eudyptes chrysocome) colony, Kidney Island, Falkland Islands, October. Vulnerable species
Gentoo penguin (Pygoscelis papua) walking back to the colony, Sea Lion Island, Falkland Islands, October
Mute swan (Cygnus olor) cygnets with head submerged while feeding underwater, with mother watching and helping, Derbyshire, England, UK. June
Eurasian griffon vulture (Gyps fulvus) in flight, Cevennes, France, March 2016
Madagascar / Malagasy kingfisher (Alcedo vintsioides) perching, Anjajavy Private Reserve, north west Madagascar
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
Adult King Vulture (Sarcoramphus papa) Laguna del Lagarto, Boca Tapada, Caribbean slope, Costa Rica, Central America
Kori bustard (Ardeotis kori) male courtship display, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Bluethroat (Luscinia svecica) singing on a branch Vendee, France, May
Seychelles Sunbird (Cinnyris dussumieri), adult male singing, Praslin Island, Republic of Seychelles
Starling (Sturnus vulgaris), eating fruit of Japanese Persimmon (Diospyros kaki), Provence, France
Brown booby (Sula leucogaster), adult diving for fish, Los Roques, Venezuela
Guianan cock-of-the-rock (Rupicola rupicola), two males fighting during lek, Guianan Shield, Suriname
Southern elephant seal (Mirounga leonina), fight between two males, Saint Andrew, South Georgia
Series of Bald eagle (Haliaeetus leucocephalus) in flight, Homer, Alaska, USA, March. Digital composite
