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Galapagos penguin (Spheniscus mendiculus), pair courting, Isabela Island, Galapagos Islands
Galapagos hawk (Buteo galapagoensis), juvenile plumage, Bainbridge Crater, Galapagos Islands
Darwins cactus finch (Geospiza scandens), feeding on Opuntia cactus flower, Santa Cruz Island, Galapagos Islands
Kingfisher ( Alcedo atthis) male and female perched on a root when building the nest, Lorraine, France, March
RF - Atlantic puffin (Fratercula arctica) flying, Hornoya, Varanger, Finnmark, Norway (This image may be licensed either as rights managed or royalty free.)
Pied avocet (Recurvirostra avosetta) adult parent and chick together. Suffolk, UK. June
Mistle thrush (Turdus viscivorus) calling perched on rowan tree. London, UK. December
Satin bowerbird (Ptilonorhynchus violaceus), maintaining his bower, Ravenshow, North Queensland, Australia, October
Golden Bowerbird (Prionodura newtoniana), placing flowers on bower, Ravenshoe, North Queensland, Australia, October
Eastern rockhopper penguins (Eudyptes chrysocome filholi), Auckland Island, December
Crested tit (Lophophanes cristatus) silhouette, Sierra de Grazalema Natural Park, Cadiz, Andalusia, Southern Spain. October
Rowi or Okarito Brown Kiwi (Apteryx rowi) male called Scooter in rainforest habitat. Okarito Forest, Westland, South Island, New Zealand
Mute swan (Cygnus olor) in mist, Bolam Lake Country Park, Northumberland, UK, November
Aerial view of green algae and Lesser flamingos (Phoeniconaias minor) in flight at the mouth of a river, Lake Bogoria, Kenya
Common swift (Apus apus) flying low to water while hunting, Norfolk, England, UK, July. Winner of Animal Behaviour Category of the British Wildlife Photography Awards 2019. Photographer
White-tailed sea eagle (Haliaeetus albicilla) flying over water with male Common eider prey (Somateria mollissima), Uto, Parainen, Finland, April
Male Black grouse (Tetrao / Lyrurus tetrix) in flight. Kuusamo, Finland, April
Laysan albatross (Phoebastria immutabilis) chick in the nest, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, February
Buzzard (Buteo buteo) landing on branch, winter, Lorraine, France, February
Boat-tailed grackle pair (Quiscalus major) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Aerial view of Griffon vulture (Gyps fulvus) flock scavenging dead pig, Cantabria, Spain. January
Ruby-throated hummingbird (Archilochus colubris) male landing on Virginia creeper (Parthenocissus quinquefolia). Hill Country, Texas, USA
Kingfisher (Alcedo atthis) male, after diving, taking off from water with fish, a Common Roach (Rutilus rutilus) Lorraine, France, July
White-tailed eagle (Haliaeetus albicilla) flying over water with fish in talons. Norway. August
Great grey owl (Strix nebulosa) in flight in woodland during snowfall. Kuusamo. Finland. February
Barred Owl (Strix varia) fledglings. Washington County, Oregon, USA. June
Griffon vulture (Gyps fulvus) soaring over Los Mallos de Riglos, Huesca province, Aragon, Spain
African fish eagle (Haliaeetus vocifer) swoops to catch a freshly caught fish, dropped by a Saddle-billed stork (Ephippiorhynchus senegalensis) after being pressurised to do so by the eagle
Taiwan Blue Magpie, (Urocissa caerulea) endemic species, Taipei city park, Taiwan
Malayan night heron (Gorsachius melanolophus) Taipei city park, Taiwan
Black-faced Spoonbill, (Platalea minor), Tainan, Taiwan. Endangered species
Taiwan blue magpie, (Urocissa caerulea), in Taipeh, Taiwan, endemic species
Taiwan yuhina (Yuhina brunneiceps) perched amongst pink blossom, Alishan National Scenic Area, Taiwan. Endemic species
Taiwan blue magpie, (Urocissa caerulea) flying, in Taipeh, Taiwan, endemic species
Glossy ibis (Plegadis falcinellus) in marshland. Donana National Park, Spain
Mute swan (Cygnus olor) stretching its wings after preening on a misty morning. Valkenhorst Nature reserve, Valkenswaard, The Netherlands, May
Mikado pheasant (Syrmaticus mikado) male, close up photo, Taiwan
Bittern (Botaurus stellaris) with marsh frog that has just woken up from hibernation, Kiskunsag National Park, Hungary
Lawess parotia (Parotia lawesii) male displaying. Papua New Guinea
Peregrine falcon (Falco peregrinus) in flight with open beak. The Netherlands. May
Crested drongo, (Dicrurus forficatus), Berenty National Park, Madagascar
Stellers sea eagle, (Haliaeetus pelagicus), on sea pack ice, Okhotsk Sea, Rausu, Hokkaido, Japan
California condor (Gymnogyps californianus) landing, forest in background. Bird with tags on wings. Near San Pedro Martir National Park, Northern Baja California, Mexico. 2017
Turtle dove (Streptopelia turtur) pair, Teide National Park, Tenerife, Canary Islands, Spain, July
Palm Cockatoo (Probosciger aterrimus) feeding on Beach Almond (Terminalia catappa) Lockhart River Foreshore, Lockhart River, Cape York Peninsula, Australia
Rufous-tailed hummingbird (Amazilia tzacatl) feeding from Heliconia flower (Heliconia champneiana) La Selva Field Station, Costa Rica
RF - Red-and-green macaws (Ara chloropterus) two with one taking off, Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
European bee eaters (Merops apiaster) one taking off another perched, Kiskunsagi National Park, Pusztaszer, Hungary. May
