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Manchurian crane (Grus japonensis) pair in courtship dance during snowstorm. Hokkaido, Japan. March
African penguin (Spheniscus demersus) standing on rock at dusk, with clustered penguins in background. South Africa. November
Greater flamingo (Phoenicopterus roseus) pair at rest in water, Cape Town, South Africa, July
Atlantic puffin (Fratercula arctica) swimming on the ocean surface, Svalbard, Norway, June
RF - Greater flamingo (Phoenicopterus roseus) flock blurred, Camargue, France
Greater flamingo (Phoenicopterus roseus) legs of bird in flight, Pont Du Gau Park, Camargue, France
Eared grebes (Podiceps nigricollis) stand erect in water during courtship, North Park, Colorado, USA. June
Grey peacock-pheasant (Polyplectron bicalcaratum) Tongbiguan Nature Reserve, Dehong, Yunnan, China
Temmincks tragopan pheasant (Tragopan temminckii) Chongqing region, Sichuan, China
Golden pheasant (Chrysolophus pictus) male and female, Yangxian nature reserve, Shaanxi, China
Kalij pheasant (Lophura leucomelanos) Tongbiguan Nature Reserve, Dehong, Yunnan, China
Scopolis shearwater (Calonectris diomedea) in flight. Tenerife, Canary Islands
Brunnichs guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018
Southern greyheaded sparrows (Passer diffusus) reflected in waterhole, Zimanga private game reserve, KwaZulu-Natal, South Africa
Pied crow (Corvus albus) pulling tail feather of whitebacked vulture, Zimanga private game reserve, KwaZulu-Natal, South Africa
Magnificent frigatebird (Fregata magnificens) in flight over sea between Isabela Island and Fernandina Island, Galapagos
Mute swan (Cygnus olor), two cygnets sheltering under parent's wing. Richmond Park, London, England, UK. April
Mute swan (Cygnus olor) swimming with cygnets in misty lake in morning light. Richmond Park, London, England, UK. May. Cropped image
Mute swan (Cygnus olor) cygnet swimming, cocking head. Barnes, London, England, UK. April
Coscoroba swan (Coscoroba coscoroba) at nest with eggs. La Pampa, Patagonia, Argentina. February
Greater rhea (Rhea americana) nest with many eggs, at edge of arable field. Patagonia, Argentina. October
California condor (Gymnogyps californianus) in flight, wings radio tagged. California condor recovery program, Sierra de San Pedro Martir National Park, Baja California Peninsula, Mexico. 2011
Pesquets parrot (Psittrichas fulgidus) perched on branch. Beauval Zoo Parc, France. Captive
Blue-faced honeyeater (Entomyzon cyanotis) perched on bark. Nitmiluk National Park, Northern Territory, Australia
Red-collared lorikeet (Trichoglossus rubritorquis) head peering out of nest box.Territory Wildlife Park, Darwin, Northern Territory, Australia. Captive
Austral blackbirds (Curaeus curaeus) and Austral thrush (Turdus falcklandii) nectaring on Blue puya (Puya berteroniana). Pollen transferred to birds heads during feeding. Parque Ingles, Chile
Southern double-collared sunbird (Cinnyris chalybeus), male nectaring on Pincushion protea (Leucospermum cordifolium). Cape, South Africa. August
Red wattlebird (Anthochaera carunculata) nectaring on Grevillea (Grevillea preissii). Western Australia Botanic Garden, Kings Park, Perth, Australia. November
Malachite sunbird (Nectarinia famosa) male nectaring on King protea (Protea cynaroides). Cape Province, South Africa. August
Purple throated mountain gem hummingbird (Lampornis calolaemus) hovering as it nectars on Porterweed (Stachytarpheta frantzii). Costa Rica
Woodland Kingfisher (Halcyon senegalensis) perched in tree, Serengeti, Tanzania
Willow ptarmigan (Lagopus lagopus) resting in snow, Taymyr Peninsula, Siberia, Russia. March
Snowy owl (Bubo scandiacus) Female taking off the winter tree, Quebec, Canada
Crested tit (Lophophanes cristatus) perched above pine forest with a series of small lochans in background, Cairngorms National Park, Scotland, UK, December 2014
Trumpeter hornbill (Bycanistes bucinator) feeding, captive
RF - Rainbow lorikeet (Trichoglossus moluccanus) portrait, captive, with digitally added leaf pattern
Little penguin (Eudyptula minor) standing on rock, silhouetted by Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016
Great horned owl (Bubo virginianus) preening two chicks in nest in Saguaro (Carnegiea gigantea) cactus, Sonoran desert, Arizona, USA
Little penguin (Eudyptula minor) standing on rocks of St Kilda breakwater, returning to nest at dusk. Melbourne city in background. Victoria, Australia. December 2016
Hoatzin (Ophisthocomus hoazin) perched on wooden bar overlooking oxbow lake. Manu National Park, Madre de Dios, Peru. September
Wandering albatross (Diomedea exulans) flying on the open ocean, Drake passage, Antarctic Peninsula, Antarctica
Gentoo penguins (Pygoscelis papua) walking in line, returning to nesting area, Port Charcot, Antarctic Peninsula, Antarctica
Little Owl (Athene noctua) immature perched in oak tree, London. July
Barn owl (Tyto alba) hovering whilst hunting in flight. Suffolk, UK. June, Cropped
Scarlet Macaws (Ara macao) drinking water inside an Amazonian claylick. Yasuni National Park, Orellana, Ecuador
Indian peafowl or blue peafowl (Pavo cristatus) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India
Northern ravens (Corvus corax) interacting in snow, Leon, Spain, February
Eurasian blue tit (Cyanistes caeruleus) on a branch, Vendee, France, January
