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Hourglass treefrog (Dendropsophus ebraccatus) male with inflated vocal sac calling at night, Central Caribbean foothills, Costa Rica
Marine iguanas (Amblyrhynchus cristatus) basking on volcanic rock, Punta Espinosa, Fernandina Island, Galapagos Islands, East Pacific Ocean
Leafcutter ants (Atta sp) colony harvesting a banana leaf, Costa Rica. 3rd place in the Insects and Spiders category of the Terre Sauvage Nature Images Awards competitions 2015
Great Green Macaw (Ara ambiguus) close up of feathers, El Manantial Macaw Sanctuary, Costa Rica. Captive
Tropical forest, Andasibe-Mantadia National Park, Madagascar, March 2005
Detail of palm leaf, backlit in sunshine, New Caledonia
Morpho butterfly (Morpho peleides) photographed on white background. Costa Rica, March
RF- Elongate surgeonfish (Acanthurus mata) school. Indonesia
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Atlantic bluefin tuna (Thunnus thynnus) within tuna farm, containing around 1000 per net. Saint Pauls Bay, Malta. Mediterranean Sea
Close up of pattern of adult Emperor angelfish (Pomacanthus imperator) Tulamben, Bali, Indonesia. Java Sea
Mirrored portrait of a Weedy scorpionfish (Rhinopias frondosa). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea. Digitally manipulated (the face is reflected down mid-line)
Longnose hawkfish (Oxycirrhites typus) hiding in coral, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
RF - Long-spined sea urchin (Astropyga radiata). Lembeh Strait, North Sulawesi, Indonesia
Scarlet macaw (Ara macao) flying, blurred motion. Costa Rica
Verreauxs sifaka (Propithecus verreauxi) running across a road with bus in background, Berenty Private Reserve, southern Madagascar, August 2016
Mother and baby Bornean orangutan (Pongo pygmaeus) in trees Tanjung Puting National Park, Borneo-Kalimatan, Indonesia, endangered species
Hoffmanns Two-toed Sloth (Choloepus hoffmanni) orphaned babies in rehabilitation program, Aviarios Sloth Sanctuary, Costa Rica
Asiatic elephants (Elephas maximus indicus) sniffing the air as they get close to a water body, Kaziranga National Park, Assam, India
Female Tassled wobbegong shark (Eucrossorhinus dasypogon) rests alongside stand of Barrel sponges (Xestospongia testudinaria) Blue Magic, Raja Ampat, West Papua, Indonesia
Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December
Adult female Fosa (Crytoprocta ferox) stalking prey on deciduous forest floor. Kirindy Forest, western Madagascar. November 2010
Galapagos green turtle (Chelonia agassizii) surfaces to breathe. Floreana Island, Galapagos Islands, Ecuador. East Pacific Ocean
Coquerels sifaka (Propithecus coquereli) Palmarium, Atsinanana, Madagascar
Indri (Indri indri) feeding on fresh leaves / shoots in the rainforest canopy. Andasibe-Mantadia National Park, eastern Madagascar
Highland cloud forest in the Talamanca Mountains of Costa Rica, April 2015
RF - Granular poison frog (Oophaga granulifera). Osa Peninsula, Costa Rica. Vulnerable species
RF - Lowland rainforest, Osa Peninsula, Costa Rica
Cup fungus (Cookeina sp) growing on decaying wood on the rainforest floor, Corcovado National Park, Osa Peninsula, Costa Rica
Aerial view of Delta Sierpe River Terraba, Corcovado National Park, Osa Peninsula, Puntarenas Province, Costa Rica. December 2014
Yellow-eyed Penguin (Megadyptes antipodes) walking to nest hidden inland in dense vegetation. Sandy Bay, Enderby Island, Auckland Group, New Zealand Sub Antarctic Islands
Blue sheep (Pseudois nayaur) Hemis National Park, Jammu and Kashmir, India
Feral domestic rabbit (Oryctolagus cuniculus) juvenile running with dead leaf in mouth, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Booby (Sula nebouxii) looking at camera underwater, Galapagos Island
Parachute daisies (Ursinia anthemoides) Little Karoo, Western Cape, South Africa
Japanese macaque (Macaca fuscata fuscata) mother with rare white furred baby, Jigokudani Valley, Nagano Prefecture, Japan. June
Verreauxs sifaka (Propithecus verreauxi) female carrying baby, Berenty Private Reserve, Madagascar. Endangered species
Feral domestic rabbit (Oryctolagus cuniculus) Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Las Gralarias Glassfrog (Nymphargus lasgralarias) spawn with tadpoles developing in eggs hanging from leaf, Mindo, Pichincha, Ecuador
Pinocchio lizard (Anolis proboscis) male on stem, Mindo, Pichincha, Ecuador, Endangered species
Two Limon glass frogs (Sachatamia ilex) sitting on plant, one above the other, Canande, Esmeraldas
Minute Glassfrog (Centrolene peristictum) male guarding eggs, Mindo, Pichincha, Ecuador, Vulnerable species
Juvenile Mouse opossum (Marmosops impavidus) feeding on moth, Pinas, El Oro, Ecuador
Mountain gorilla (Gorilla beringei beringei) female carrying baby on her back, member of Humba group, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Close-up of an eye of a Sumatran tiger (Panthera tigris sumatrae), captive, occurs in Sumatra, Indonesia
RF- Juvenile Sumatran tiger (Panthera tigris sumatrae), aged four months, captive, occurs in Sumatra, Indonesia
Japanese red squirrel (Sciurus vulgaris orientis) struggling with long branch collected for construction of nest. Hokkaido, Japan
