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Blue whale (Balaenoptera musculus) diving beneath ocean surface. Indian Ocean, Sri Lanka
Blue-footed booby (Sula nebouxii) courting pair, South coast, Santa Cruz Island, Galapagos
Sinbad the male Kakapo (Strigops habroptilus) curiously peering from the bushes during the day. Codfish Island, Stewart Island, New Zealand, January. Critically endangered
Leaf cutter ants (Atta sp) carrying colourful plant matter, reflected in water, Laguna del Lagarto, Santa Rita, Costa Rica
Leaf cutter ants (Atta sp) carrying plant matter, Costa Rica
Santa Cruz Galapagos tortoise (Chelonoidis nigra porteri) feeding on grass, Santa Cruz Highlands, Galapagos
Male Leach's sea star (Leiaster leachi) broadcast spawning, releasing streams of sperm from its arms. Kagoshima Prefecture, Kyushu, Japan
RF - Ocellate octopus (Amphioctopus mototi) swimming up from the sea floor. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Splendid leaf frog (Cruziohyla calcarifer) La Selva Field Station, Costa Rica
Siberian flying squirrel (Pteromys volans orii) flying directly over photographer. Hokkaido, Japan. February
Indo-Pacific bottlenose dolphin (Tursiops aduncus) with penis extended. Ogasawara / Bonin Islands, Japan
Marine iguana (Amblyrhynchus cristatus) swimming underwater, Fernandina Island, Galapagos
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
RF - Keel-billed toucan (Ramphastos sulfuratus) perched on mossy branch. Boca Tapada, Costa Rica. (This image may be licensed either as rights managed or royalty free.)
Male Fossa (Cryptoprocta ferox) climbing down tree trunk from forest canopy. Mid-altitude rainforest, Andasibe-Mantadia National Park, eastern Madagascar. Endangered
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
Stellers sea-eagle (Haliaeetus pelagicus) in flight, Hokkaido, Japan, February
Japanese macaque (Macaca fuscata) sleeping at hot spring in Jigokudani, Yaenkoen, Nagano, Japan, February
Galapagos penguin (Spheniscus mendiculus) underwater, Bartholome Island, Galapagos Islands, East Pacific Ocean
Snow leopard (Panthera uncia) female in snow, Hemis National Park, Ladakh, India
Bamboo lemur (Hapalmur griseus) in tree. Andasibe-Mantadia National Park, Madagascar
Japanese crane (Grus japonensis) displaying, wings in heart shape, Hokkiado, Japan, February
Kaleidoscopic image of Clarks anemonefish (Amphiprion clarkii), Ambon, Indonesia
White-vented storm petrel (Oceanites gracilis galapagoensis) running across water, offshore Cape Berkeley, Isabela island, Galapagos
Blue-footed booby (Sula nebouxii) landing, South coast, Santa Cruz Island, Galapagos
Great frigatebird (Fregata minor) in tree with pair of Nazca boobies (Sula granti) showing mating behaviour, Genovesa Island, Galapagos
Red-billed tropicbird (Phaethon aethereus) in flight, Plazas Island, Galapagos
Kakapo (Strigops habroptilus) close up showing sensory facial feathers, Codfish Island / Whenua Hou, Southland, New Zealand, February. Critically endangered
Australasian crested grebe pair (Podiceps cristatus australis) in courtship display. Ashburton Lakes, Canterbury, New Zealand. August
Veined octopus (Amphioctopus marginatus) resting on top of the two halves of an old clam shell on the sandy seabed. These octopus regularly carry pieces of shell so they can use them as protective
Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
RF - Mantis shrimp (Odontodactylus scyllarus) on walk about on coral reef. Puerto Galera, Philippines
Nudibranch (Phyllidia marindica), Lembeh Strait, North Sulawesi, Indonesia. December
Bornean Orangutan (Pongo pygmaeus) female face portrait, Tanjung Puting reserve, Camp Leakey, Central Kalimantan, Borneo
Whooper swans (Cygnus cygnus) three coming into land, Hokkaido, Japan, February
Swallow-tailed gull (Creagrus furcatus) Plazas Island, Galapagos
Scarlet macaws (Ara macao) La Selva, Costa Rica
Japanese cranes (Grus japonensis) displaying in snow, Hokkiado, Japan, February
Stellers sea-eagle (Haliaeetus pelagicus) landing on pack ice, Hokkaido, Japan, February
Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea
Portrait of Bigfin squid (Sepioteuthis lessoniana) hovering in open water above a coral reef at night. Dampier Strait, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Ariel toucan (Ramphastos ariel) perched on branch, Atlantic Rainforest, Rio de Janeiro city, Rio de Janeiro State, Brazil
Imbabura tree frog (Hypsiboas picturatus) portrait, Canande, Esmeraldas, Ecuador
Tropical shield mantis (Choeradodis sp. ) (Possibly C. rhombicollis) Canande, EsmeraldasTropical shield mantis (Choeradodis sp.) (Possibly C. rhombicollis) Canande, Esmeraldas, Ecuador
Portrait of Sally lightfoot crab (Grapsus grapsus) on a beach. San Cristobal Island, Galapagos Islands, Ecuador. Tropical East Pacific Ocean
Black-crowned Central American squirrel monkey (Saimiri oerstedii) pair close-up Manuel Antonio National Park, Quepos, Costa Rica
Brown pelican (Pelecanus occidentalis) close up whilst resting, Punta Suarez, Espanola Island, Galapagos