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Galapagos green turtle (Chelonia mydas agassizi) swimming in coastal waters of San Cristobal Island, Galapagos
Sergeant major (Abudefduf sp) schooling in mangroves of Isabela Island, Galapagos
Resplendent quetzal (Pharomachrus mocinno) male perched on branch. Costa Rica
Rivolis hummingbird (Eugenes fulgens) nectaring on Fuchsia (Fuchsia sp) flower. Costa Rica
Leaf-nosed bat (Phyllostomidae sp) flying towards Banana (Musa sp) flower to feed. Costa Rica
Green-crowned brilliant (Heliodoxa jacula), three feeding on Lobster-claw (Heliconia sp) flower in rain. Costa Rica
Malachite sunbird (Nectarinia famosa) feding at flower, Kirstenbosch National Botanical Garden, Cape Town, South Africa, September
Orange-breasted sunbird (Anthobaphes violacea) female on Leucospermum cordifolium Yellow Bird Kirstenbosch National Botanical Garden, Cape Town, South Africa, September
Cape sugarbird (Promerops cafer) on king protea, Kirstenbosch National Botanical Garden, Cape Town, South Africa, September
Southern double-collared sunbird (Cinnyris chalybeus), Kirstenbosch National Botanical Garden, Cape Town, South Africa, September
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Flightless cormorant (Phalacrocorax harrisi), adult and two chicks. Beagle Crater, Isabela Island, Galapagos
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Himalayan brown bear (Ursus arctos isabellinus) female with two young cubs climbing up snowy slope. Western Ladakh, northern India
Bamboo lemur (Hapalmur griseus) sitting on tree stump. Andasibe-Mantadia National Park, Madagascar
Snow leopard (Panthera uncia) male running hunting Himalayan ibex on the snow in Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India, February
Himalayan ibex (Capra sibirica) male on snow, Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India, February
Tapanuli Orangutan (Pongo tapanuliensis) Beti, juvenile female approximate age 6 years, playing with mother, Beta, Batang Toru Forest, Sumatran Orangutan Conservation Project
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Montezuma oropendola (Psarocolius montezuma) hanging upside down as part of a courtship display. Costa Rica
Double-crested basilisk (Basiliscus plumifrons) running across water surface, Santa Rita, Costa Rica
Yellow warty frogfish (Antennarius maculatus) Seraya, Tulamben area, Bali, Indonesia. Java Sea
Galapagos racer snakes (Pseudalsophis biserialis) group alert watching for prey on beach, Galapagos, June. Endangered species. Taken on location for BBC tv Blue Planet 2 series
Mountain landscape reflected in lake, Basongcuo National Park, Qinghai-Tibet Plateau, Tibet, November 2011
Adult Kea (Nestor notabilis) close-up portrait. Arthurs Pass National Park, New Zealand, July. Endangered Species
Bornean orangutan (Pongo pygmaeus) male, Gunung Palung National Park, Borneo
Leaf mimic bush cricket or katydid (unknown species, family Tettigoniidae) in the middle of moulting its skin. Moulting from one sub-adult stage (instar) to another is known as ecdysis
White fronted capuchin monkey (Cebus capucinus) feeding, March, Costa Rica
Madagascar / Malagasy kingfisher (Alcedo vintsioides) perching, Anjajavy Private Reserve, north west Madagascar
Eastern Ring-tailed Vontsira (Galidia elegans elegans) (endemic family Eupleridae) foraging on forest floor. Marojejy National Park, north east Madagascar
Adult King Vulture (Sarcoramphus papa) Laguna del Lagarto, Boca Tapada, Caribbean slope, Costa Rica, Central America
Female Red-eyed Tree Frog (Agalychnis callidryas) - Caribbean slope race (blue flanks) - obscured in rainforest foliage. Mid-altitude rainforest near Aranal, central Costa Rica
White-headed capuchin (Cebus capucinus) walking on giant palm-tree, Costa Rica
Sea cucumber / Sea apple (Pseudocolochirus violaceus) Rinca, Komodo National Park, Indonesia
Galapagos giant tortoise (Geochelone nigra porteri), Cerro El Chato, Santa Cruz Island, Galapagos Islands
Brazilian tapir (Tapirus terrestris) portrait, Tambopata Research Centre, Peru
Komodo dragon (Varanus komodoensis) Rinca Island, Indonesia
Sea apple (Pseudocolochirus violaceus) Komodo National Park, Indonesia
Masked tree frog (Smilisca phaeota) El Arenal region, Costa Rica
Toque Macaque (Macaca sinica) baby peering through mothers legs, Yala National Park, Southern Province, Sri Lanka
Sumatran orangutan (Pongo abelii) female next to infant, Gunung Leuser National Park, Indonesia, UNESCO World Heritage site, November
Sumatran orangutan (Pongo abelii) female moving between lianas, Gunung Leuser National Park, UNESCO World Heritage site, November
Rolling farmland in the Overberg region near Villiersdorp, Western Cape, South Africa. December 2014
Lowland Streaked Tenrec (Hemicentetes semispinosus) active on forest floor at night. Masoala National Park, north east Madagascar
Mandarinfish (Synchiropus splendidus) Yap, Micronesia
Malabar pit viper (Trimeresurus malabaricus), green colour morph. Agumbe, Karnataka, Western Ghats, India. Endemic
King cobra (Ophiophagus hannah), low wide angle perspective Agumbe, Karnataka, Western Ghats, India. Vulnerable species
Proboscis bats (Rhynchonycteris naso) day roosting, Costa Rica, March 2014
