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Yellow-flecked glassfrogs (Sachatamia albomaculata) mating pair, Osa Peninsula, Costa Rica
Veragua cross-banded treefrogs (Smilisca sordida) mating pair at night, Osa Peninsula, Costa Rica
Green mandarinfish (Synchiropus splendidus) pair courting and mating at sunset, Sulu Sea, Philippines
Sarayacu clownfrog (Dendropsophus sarayacuensis) with its clutch of eggs developing and starting to drop off from a leaf into the water below, where the tadpoles will develop
Humpback whale (Megaptera novaeangliae), with flipper raised in the air above water, Machalilla National Park, Manabi, Ecuador
Cat-eyed snake (Leptodeira septentrionalis) predating Olive snouted treefrog (Scinax elaechroa) in Tortuguero National Park, Costa Rica. November 2013
Redshank (Tringa totanus) silhouetted, Iceland, June
Red-winged blackbird (Agelaius phoeniceus) male in territorial display, Virginia, USA. May
Humpback whale (Megaptera novaeangliae) blowing, Prince William Sound, Alaska, USA, July
Saturniid moth (Saturniidae) with wings open to reveal eyespots, a means of deterring predators, Caribbean Slopes, Cordilera de Talamanca, Costa Rica
Golden eagle (Aquila chrysaetos) male head portrait beak open calling, Kalvtrask, Vasterbotten, Sweden. December
Brydes whale (Balaenoptera brydei) breaching, Raja Ampat, Western Papua, Indonesian New Guinea
Green turtle (Chelonia mydas) female digging nest, North coast, Santa Cruz Island, Galapagos
Alcedo giant tortoise (Chelonoidis vandenburghi) group resting in pool, Alcedo Volcano, Isabela Island, Galapagos
Ring-tailed cardinalfish (Ostorhinchus aureus), male protecting and incubating eggs in mouth, Philippines
Pink-toed tarantula (Avicularia avicularia), female waiting in ambush outside daytime lair in palm tree trunk. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
RF - Pink anemonefish (Amphiprion perideraion) with host anemone (Heteractis magnifica). Bunaken National Park, North Sulawesi, Indonesia
Golden jackal (Canis aureus) howling in grassland. Danube Delta, Romania, May
African wild dog (Lycaon pictus), male and female play fighting. Mana Pools National Park, Zimbabwe
Money spider (Tenuiphantes sp) tiptoeing behaviour on dandelion seed head. Spiders can stand on tiptoes and produced silk to balloon and travel with the airflow
Corn bunting (Emberiza calandra) singing, Prado del rey, Cadiz, Spain, March
Spotted nutcrackers (Nucifraga caryocatactes) fighting in snow, Vitosha Mountain, Sofia, Bulgaria
Indo-Pacific humpback dolphins (Sousa chinensis) at surface, Tai O, Lantau Island, Hong Kong, China
Black egret (Egretta ardesiaca) forming wings into umbrella whilst fishing, Gambia
Tibetan macaque (Macaca thibetana) infant in tree, Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Jaguar (Panthera onca) four month old cub washing, native to Southern and Central America, captive
Arctic fox ( Vulpes lagopus) in snow carrying snowgoose egg, Wrangel Island, Far East Russia, May
Demoiselle cranes (Anthropoides virgo) flying at sunrise during migration. Khichan, Western Rajasthan, India. December
Tibetan macaque (Macaca thibetana) juveniles playing, hanging upside down in tree, Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
White-rumped snowfinch (Montifringilla taczanowskii) two fighting, Tibetan Plateau, Qinghai, China
Eastern tiger swallowtail butterflies (Papilio glaucus) puddling, New Brunswick, Canada, June
Pumas (Puma concolor) stalking a troop of howler monkeys, Corcovado National Park, Costa Rica, May
Long-tailed manakin (Chiroxiphia linearis) perched on lek (display perch), Costa Rica
Short finned pilot whale (Globicephala macrorhynchus) about to surface, Baja California, Mexico, February
Humpback whale (Megaptera novaeangliae) breaching - leaping out of the water, Sea of Cortez, Baja California, Mexico
Whitethroat (Sylvia communis) male perched on Willow branch in spring, Wiltshire, England, UK, April
Orange tip butterfly (Anthocharis cardamines) female and male, visiting Honesty flower (Lunaria annua) Wiltshire, England, UK, May
Sooty Albatross (Phoebetria fusca) group courting on high inland ridge. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic. Endangered
Emperor shrimp (Periclemenes imperator) on sea cucumber (Thelenota ananas) Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Cheetah (Acinonyx jubatus) male on the hunt with Crater rim in the background, Ngorongoro Crater, Tanzania
American red squirrel (Tamiasciurus hudsonicus) collecting, Douglas Fir (Pseudotsuga menziesii) cones, Montana, USA. October
Yellow warbler (Dendroica petechia) collecting nesting material from Bulrush cattail (Typha sp) Bozeman, Montana. USA, June
RF - Red deer (Cervus elaphus) stags fighting, Holkham Park, Norfolk, England, UK, September
Garden Cross Spider (Araneus diadematus) wrapping its Common Carder Bee (Bombus pascuorum) prey in silk, Bristol, UK, September. Sequence 4/10
Small flock of Dunlin (Calidris alpina) with a Sanderling (Calidris alba) flying in front of crashing waves. Druridge Bay, Northumberland, England, UK, January
Fungus gnat (Mycetophilidae) larvae, with sticky hanging threads, waiting to ambush small flying insects. Deer caves, Gunung Mulu National Park, Borneo, Sarawak, Malaysia
Oriental dwarf kingfisher (Ceyx erithacus) catching and eating a fish at Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan Province, China, April
Raggy scorpionfish (Scorpaenopsis venosa) disputing territory. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
