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Whitebacked vulture (Gyps africanus) fighting over food, Zimanga private game reserve, KwaZulu-Natal, South Africa
Alcedo giant tortoise (Chelonoidis vandenburghi), group bathing. Alcedo Volcano, Isabela Island, Galapagos. 2017
Southern elephant seal (Mirounga leonina), two males, equally matched with mouths open in aggression. Out of focus King penguin (Aptenodytes patagonicus) colony in background
Red fox (Vulpes vulpes) sequence of snow diving whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February 2019. Digital composite
King penguin (Aptenodytes patagonicus), two sleeping on snow. Grytviken, South Georgia. March
Red deer (Cervus elaphus) stag bellowing during rut in grassland, doe in background
Spinner dolphin (Stenella longirostris) pod socialising close to surface. Fury Shoal, Sataya Reef, Marsa Alam, Egypt
Southern royal albatross (Diomedea epomophora) sub-adults, group displaying in tussock grassland. Campbell Island, New Zealand. February
African elephants (Loxodonta africana) herd silhouetted at sunset. Chobe National Park, Botswana
Dalmatian wall lizard, (Podarcis melisellensis), fight between two males with distinct morphs, Croatia, April. Non-ex
Rainbow lorikeet (Trichoglossus moluccanus) feeding chicks in nest hole. Lane Cove National Park, Sydney, New South Wales, Australia
Brooks urchin shrimp (Allopontonia brooki), commensal among the venomous spines of Variable sea urchin (Asthenosoma varium). On sea floor, Komodo, Indonesia
Bengal Tiger (Panthera tigris), Arrowhead and cub cooling off in water, Ranthambore National Park, India
Bengal Tiger (Panthera tigris) Arrowhead patrolling his territory, Ranthambore National Park, India
Bengal Tiger (Panthera tigris) Arrowhead chasing stag near lake sequence, Ranthambore National Park, India Sequence 7 of 7
Red admiral (Vanessa atalanta) basking in sun, on Snowdrops (Galanthus nivalis). In garden, Surrey, England, UK. February
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
Bar headed geese (Anser indicus), four in flight, one calling. Coming in to land at breeding site, Qinghai Lake, Qinghai Province, China
Amur / Siberian tiger (Panthera tigris altaica) pair in courtship, in snow. Captive in tiger park, Heilongjiang Province, China. January
Chinese white cheeked gibbon (Nomascus leucogenys) female hanging from tree, carrying baby aged two weeks. Parents released into wild from captive breeding programme
African lion, (Panthera leo) confronted by a herd of African buffalo / Cape buffalo (Syncerus caffer), Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
Egyptian goose (Alopochen aegyptiacus) bathing, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
RF - Pied kingfisher (Ceryle rudis) group of three perched on branch, squabbling, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
African Spoonbill (Platalea alba) preening, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
Great egret (Ardea alba) with prey in beak, in evening light. Myakka River State Park, Florida, USA, March. Sequence 2/2
Great egret (Ardea alba) diving head into water to catch prey, in evening light. Myakka River State Park, Florida, USA, March. Sequence 1/2
Tricoloured heron (Egretta tricolor) fishing by flying low over water, amongst Water lilies. Everglades National Park, Florida, USA. March
Three Pink anemonefish (Amphiprion perideraion) living in a purple skirted magnificent sea anemone (Heteractis magnifica) on a coral reef, below a small island
Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef
Reef lizardifsh (Synodus variegatus) at a cleaning station, being cleaned by Holthuis anemone shrimps (Ancylomenes holthuisi). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Waved albatross (Phoebastria irrorata) pair in courtship display at nest site, Punta Suarez, Espanola Island, Galapagos
Little tern (Sterna albifrons ) feeding sand eel (Hyperoplus spp) to young chick. Gronant Dunes, Denbighshire, Wales, UK, June
Black-browed albatross pair (Thalassarche melanophris) mutual grooming during copulation. New Island, Falkland Islands, November
Swallow (Hirundo rustica) feeding young on fence, Norfolk, England, UK, September
Red deer (Cervus elaphus) herd of female does and young crossing water, Jura, Scotland, UK, September
Red deer (Cervus elaphus) female does and young crossing water, Jura, Scotland, UK, September
RF - Two Red foxes (Vulpes vulpes) on hind legs play fighting, Netherlands
Yellow-rumped cacique (Cacicus cela) calling, Madidi NP, Bolivia
Red Fox (Vulpes Vulpes) cubs play fighting North London, England, UK, June
Canada geese (Branta canadensis) two during territorial fight in early spring, Ithaca. New York, USA
Snow leopard (Panthera uncia) hunting and stalking through the mountains in Serxu, Garze Prefecture, Sichuan Province, China
Black erget (Egretta gularis) displaying, Joao Vieira Poillo, Bijagos Archipellago. Guinea Bissau
Nurse sharks (Ginglymostoma cirratum) three in a courtship dance at sunrise in a mangrove area near Eleuthera, Bahamas
Common / Northern minke whale (Balaenoptera acutorostrata) surfacing, Pacific coast of Baja California, Mexico, February
Royal spoonbill (Platalea regia) preening and shaking head feathers, Victoria, Australia
Two Buzzards (Buteo buteo) fighting, Lake Csaj, Kiskunsagi National Park, Hungary. February
Cormorants (Phalacrocorax carbo) squabbling over fish, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Two Burrowing owls (Athene cunicularia) fledgling with adult about to take off, Pantanal, Brazil
