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Goosander (Mergus merganser) chick hunting mayfly, Upper Bavaria, Germany
Village weaver (Ploceus cucullatus) male bird building nest, Masai-Mara Game Reserve, Kenya
Greylag geese (Anser anser) fighting, Antwerpen, Belgium, March
Sea cucumber (Bohadschia argus) which has ejected a part of its internal organs called Cuvierian tubules. These significantly sticky strings are a defense mechanism designed to deter predators
Peringueys / Sidewinding adder (Bitis peringueyi) hiding in shallow sand, Namib Desert, Namibia. This little endemic snake is one of the smallest adders in the world
Pearlfish (Carapus acus) outside of host Sea cucumber (Stichopus regalis) at night. These species have a commensal relationship, with the fish living in the gut of the sea cucumber. Catalonia, Spain
Short eared owl (Asio flammeus) tilting head upside down and backwards, Breton Marsh, West France, June
Dingo (Canis lupus dingo) fighting on a beach. Fraser Island UNESCO World Heritage Site. Queensland, Australia, November
Gunnison sage-grouse (Centrocercus minimus) male displaying at a lek. Gunnison County, Colorado. April
RF- Capercaillie (Tetrao urogallus) male displaying, Inshriach, Cairngorms National Park, Scotland, UK. February
Tabby Persian-cross kitten, age 7 weeks, looking startled
Whooper swan (Cygnus cygnus) pair on loch in heart shape, Cairngorms National Park, Scotland, UK, November
Pine marten (Martes martes) leaping, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Animal Behaviour category of the BWPA (British Wildlife Photography Awards) Competition 2016
Humpback Whale (Megaptera novaeangliae) breaching, Machalilla National Park, Manabi, Ecuador
RF - Bottlenose dolphins (Tursiops truncatus) porpoising, Chanonry Point, Moray Firth, Highlands, Scotland. August
RF- Eagle owl (Bubo bubo) calling, raised in captivity and newly released into the wild. Stockholm archipelago, Uppland, Sweden. September
Silk moth (Automeris zugana) sequence 2 of 2, with wings open to reveal eyespots, a means of deterring predators, photographed on a white background in mobile field studio
Grass snake (Natrix natrix) flicking tongue, Hampstead Heath, London, England, UK, August
Red kite (Milvus milvus) in a dive, Gigrin Farm, Rhayader, Wales, UK April
Kingfisher (Alcedo atthis) male taking off from water after diving for fish, Worcestershire, England, UK, July
Maldive anemonefish (Amphiprion nigripes) shoal swimming in the current flowing over a Magnificent sea anemone (Heteractis magnifica). Baa Atoll, Maldives. Indian Ocean
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
Orange-finned anemonefish (Amphiprion chrysopterus) guarding their recently spawned red eggs on a rock next to the host anemone. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean
Yellowhead jawfish (Opistognathus aurifrons) spitting out sand as it digs its burrow. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Juvenile Andean mouse opossum (Marmosops impavidus) eating a moth, El Oro, Ecuador
Florida manatee (Trichechus manatus latirostrus) with Blue gill sunfish (Lepomis macrochirus) cleaning it, in a freshwater spring, beneath trees. Three Sisters Spring, Crystal River, Florida, USA
Graylag gosling (Anser anser) with parent, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. April
Common yellow throat (Geothlypis trichas) male, singing, Anchorage Provincial Park, Grand Manan Island, New Brunswick, Canada, June
Humpback whale (Megaptera novaeangliae) surfacing, rostrum above surface, Bay of Fundy, New Brunswick, Canada, November
RF- Pod of Atlantic spotted dolphins (Stenella frontalis) over shallow sand bank accompanied by larger Bottlenosed dolphin (Tursiops truncatus). Sandy Ridge, Little Bahama Bank, Bahamas
Orca whale (Orcinus orca) breaching. British Columbia, Canada, Pacific Ocean
Grizzly Bear (Ursus arctos) eating a fish, Lake Clarke National Park, Alaska, August
White rhinoceros (Ceratotherium simum) charging, Mala Mala Game Reserve, South Africa
Brown hare, (Lepus europaeus), male and female displaying courting behaviour, Islay, Scotland, UK. March
Brown hare, (Lepus europaeus), males in pursuit of female that is in season, Islay, Scotland, UK. March
Jaguar (Panthera onca) female stealthily entering the river as she begins to hunt caiman. Northern Pantanal Cuiaba River, Mato Grosso, Brazil
Lions (Panthera leo) - two brothers patrolling territorial boundary at border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
RF- Adult male Serval (Felis serval) stalking prey in long grass. Serengeti National Park, Tanzania
Alpine ibex (Capra ibex) adult males fighting on a steep mountain side, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Common cranes (Grus grus) flock on migration at sunset, Montier en Der, Aube, France, November
Sedge warbler (Acrocephalus schoenobaenus) singing with bokeh effect in background, Norfolk, England, UK, May. Winner of the Birds in Environment Category of the Bird Photographer of the Year 2015
Bengal tiger (Panthera tigris) cooling down in waterhole, portrait. Ranthambore National Park, India
Common octopus (Octopus vulgaris) hunting on a reef off Eleuthera Island, Bahamas
RF - Maldive anemonefish (Amphiprion nigripes) swimming above their host Magnificent sea anemone (Heteractis magnifica) on coral reef
RF - American flamingo (Phoenicopterus ruber) preening feathers. Captive. (This image may be licensed either as rights managed or royalty free.)
Clearfin lionfish (Pterois radiata) mating. The female (left) has just released a raft of eggs, which is visible between her tail and the red face of the male
Reef manta (Mobula alfredi) visiting a cleaning station on a coral reef with a school of Onespot snappers (Lutjanus monostigma)
Portrait of a Blackbar soldierfish (Myripristis jacobus) infected with Cymothoid isopod parasites. The large parasite between the eyes is a female, the smaller individuals are males
