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Bottlenosed dolphin (Tursiops truncatus) porpoising during annual sardine run, Port St Johns, South Africa. June
Angolan giraffes (Giraffa camelopardalis angolensis) and black rhinoceros (Diceros bicornis) reflecting in waterhole at sunset, Okaukuejo, Etosha National Park, Namibia, Africa
Nilgai or Blue bull (Boselaphus tragocamelus), silhouette of male at sunset, with Black drongo bird on its back. Velavadar National Park, Gujarat, India
Humayuns night frog (nyctibatrachus humayuni), male guarding cluster of eggs on leaf. Western Ghats Amboli, Maharashtra, India. Endemic
Mountain landscape at Tangjiahe National Nature Reserve, Sichuan Province, China April, 2015
Spoonbill (Platalea leucorodia) carrying nest material, The Netherlands
Ripe Barley (Hordeum vulgare) ears in field ready for harvest and the rising sun, Stone Henge, Salisbury, England, UK, August
Borage flower bud (Borago officinalis) near Nice, south of France, June
Peruvian mantis (Oxyopsis peruviana) silhouette with sun behind, captive, occurs in South America
Groot Schietveld at sunrise, Wuustwezel, Belgium, May 2010
Scots pine trees (Pinus sylvestris) on island in wetlands, Klein Schietveld, Brasschaat, Belgium
Boababs (Adansonia grandidieri) silhouetted at dawn, Allee des Baobabs / Avenue of the Baobabs, Morondave, Madagascar
RF - Reticulated Glass Frog (Hyalinobatrachium valerioi) backlit showing highly translucent body. Osa Peninsula, Costa Rica
Pheasant (Phasianus colchicus) male flying to roost at sunset with trees silhouetted in background England, UK. December
Dingo (Canis lupus dingo) fighting on a beach. Fraser Island UNESCO World Heritage Site. Queensland, Australia, November
RF - Grizzly Bear (Ursus arctos) at dawn, Lake Clarke National Park, Alaska, September
RF- White storks (Ciconia ciconia) nesting on poles erected by the city of Caceres in Extremadura, Spain, to compensate for the removal of an old farmhouse supporting a colony of storks
Frog spawn {Rana temporaria} just a 1-2 days old, backlit on lightbox, Cornwall, UK. January 2012
Black kitten in garden, Germany
Mute swan (Cygnus olor) at sunrise silhouetted in waters of Lamlash Bay, Isle of Arran, South Arran Marine Protected Area, Scotland, UK, August
Sea stack and Talisker Beach, Isle of Skye, Inner Hebrides, Scotland, UK, September 2013
View from Isle of Skye across Sound of Rsay to Rona and Torridon Hills, at dawn, Inner Hebrides, Scotland, UK, April 2014
Masai giraffe (Giraffa camelopardalis tippelskirchi) herd walking, Masai-Mara Game Reserve, Kenya. October
Fallow deer (Dama dama) buck crossing road under street lights. London, UK. January
Wood sandpiper (Tringa glareola) at dusk silhouetted against setting sun, Karigasniemi, Northern Finland, May. Bookplate from Danny Greens The Long Journey North
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Lion cub (Panthera leo) silhouetted at sunrise, taken on location for Pride tv series. September 2003
RF- Looking up under the gills of toadstools of Porcelain fungus (Oudemansiella mucida), Cornwall, UK
English oak tree (Quercus robur) at sunset, Monmouthshire Wales UK, March
Manta ray (Manta birostris), San Benedicto, Revillagigedo (Socorro) Islands, Mexico, East Pacific Ocean
Common Swallowtail butterfly (Papilio machaon) backlit showing wing structure. Podere Montecucco, Orvieto, Umbria, Italy, October
RF - Delicate grass and flowers silhouetted black against white background
Eagle owl (Bubo bubo) in flight through forest, backlit at dawn, Czech Republic, November. Captive
European Cave Spider (Meta menardi) in limestone cave. Plitvice Lakes National Park, Croatia. January
RF- Little blue heron (Egretta caerulea) and Great egret (Ardea alba) in lagoon at sunrise. Fort Myers beach, Gulf Coast, Florida, USA, March
Little blue heron (Egretta caerulea) in lagoon at sunrise, Fort Myers Beach, Gulf Coast, Florida, USA, March
Great white egret (Ardea alba) silhouetted at sunset, Lake Csaj, Pusztaszer, Hungary, February. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Hedge Parsley seed head (Torilis japonica) silhouetted at sunset, Norfolk, UK, November
Polar bear (Ursus maritimus) silhouetted at waters edge at sunset, Bernard Spit, off the 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, September. Vulnerable species
Common / Eurasian cranes (Grus grus) taking flight for roasting site, at sunset, silhouetted. Lake Hornborga, Sweden, April
Kelp gull (Larus dominicanus) on rocky shore with crashing waves at sunset, Bird Island, Lamberts Bay, Western Cape province, South Africa, September 2012
Two Great egrets (Ardea alba) wading, silhouetted at dawn, Keoladeo National Park, Bharatpur, Rajasthan, India
Masai giraffes (Giraffa camelopardalis tippelskirchi), at sunrise, Masai-Mara Game Reserve, Kenya. March
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at sunrise. Kasanka National Park, Zambia
Stag beetle (Lucanus cervus) silhouetted against oak tree leaf. Elbe, Germany, June
Leafcutter ants (Atta sp) colony harvesting a banana leaf, Costa Rica. 3rd place in the Insects and Spiders category of the Terre Sauvage Nature Images Awards competitions 2015
Northern lights (Aurora borealis) glowing brightly over trees along Steese Highway, Cleary Point, outside Fairbanks, Alaska
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica