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Female Red fox (Vulpes vulpes) walking along tree trunk in heavy fog at night. Light source behind the subject creates dramatic volumetric lighting. Vertes Mountains, Hungary
White-tailed eagle (Haliaeetus albicilla) flying over water with fish in talons. Norway. August
Crested tit (Lophophanes cristatus) about to land on Elmleaf blackberry (Rubus ulmifolius) plant, Sierra de Grazalema Natural Park, Southern Spain, January
Night sky over Stone arch, called the Julia arch. Ennedi Natural and Cultural Reserve, UNESCO World Heritage Site, Chad. September 2019
Bactrian camel (Camelus bactrianus) male silhouetted at sunset, living in the wild but owned by a camel herdsman, Turpan Basin, Gobi Desert, Xinjiang, China
Red fox (Vulpes vulpes) silhouetted, running along horizon, Switzerland
Mongol driving a camel caravan of Bactrian camel (Camelus bactrianus) Bashang Grassland, Zhangjiakou, Hebei Province, Mongolia, China
Red deer (Cervus elaphus) silhoutted at dusk, Richmond Park, London, England, UK. October
Aurora Borealis above silhouetted trees, northern Finland, March
Frozen water patterns in Klein Schietveld, Brasschaat, Belgium
Silhouette of an African leopard (Panthera pardus pardus) snarling, Mkuze, South Africa. September. White Fang - this leopard bared its fangs to the hyenas threatening his prey
Silhouette of the head of a Grey-headed flying-fox (Pteropus poliocephalus) through its wings. ÔÇ¿ÔÇ¿On hot days when resting in trees, flying-foxes will spread their highly vascular wings
Man using pole to propel bamboo raft on Lijang / Li River, Karst peaks beyond. At dawn, Lijiang River Scenic Zone, Yangshuo county, Guangxi, China
Little penguin (Eudyptula minor) standing on rock, silhouetted by Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016
Beech trees (Fagus sylvatica) at night in winter, with lamp post illuminating the woodland. Felbrigg Woods, Norfolk, England, UK. January
Flock of Pink footed geese (Anser brachyrhynchus) in flight at dusk, Norfolk, England, UK, October
RF- Fallow deer (Dama dama) doe at sunrise during rut, Deer Park, Holkham, Norfolk, UK, October
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
Mountain view towards Loch Nevis overlooking Knoydart peninsula with Skye beyond. Taken from Sgurr Cos na Breachd-laoidh, Lochaber, Scotland, UK, September 2016
Red deer (Cervus elaphus) male stag silhouetted at dusk, rut season, Jura, Scotland, UK, September
Red deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK. FebruaryRed deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK.February
An Orange lacewing butterfly (Cethosia penthesilea), Cairns Botanical Gardens, Queensland, Australia. Captive. August
Green-veined white butterfly (Artogeia / Pieris napi) resting on reed in late evening light, Volehouse Moor, Devon, UK. July
Abstract patterns of reflected sky and clouds on the ocean with a silhouetted Red-faced cormorant (Phalarocorax urile) in flight, St. Paul Island, Pribilof Islands, Alaska, USA, July
Great Egret (Ardea alba), about to land in water, backlit, Bolsa Chica Ecological Reserve, California, USA, October
Mixed Blackbird flock, mostly Red-winged Blackbirds (Agelaius phoeniceus), in flight against backdrop of autumn foliage in November, Bosque Del Apache National Wildlife Refuge, New Mexico, USA
European otter (Lutra lutra) group with one shaking off water, Kiskunsagi National Park, Hungary, October
European otter (Lutra lutra) two at waters edge, one shaking off water, Kiskunsagi National Park, Pusztaszer, Hungary, October
European otter (Lutra lutra) shaking off water, Kiskunsagi National Park, Hungary, January
Wood warbler (Phylloscopus sibilatrix) singing whilst perched on branch, silhouetted at dawn. Sheffield, England, UK. May
Red fox (Vulpes vulpes) silhouetted, Sado Estuary, Portugal. September
Synchronous fire flies (Pteroptyx sp. ) mangroves along the Kinabatangan River, Sabah, BorneoSynchronous fire flies (Pteroptyx sp.) mangroves along the Kinabatangan River, Sabah, Borneo
Fallow deer (Dama dama) at night, Gyulaj, Hungary Third place in the Nature Portfolio category of the World Press Photo Awards 2017
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
Sumatran orangutan (Pongo abelii) female moving between lianas, Gunung Leuser National Park, UNESCO World Heritage site, November
Aerial view of two Gemsbok (Oryx gazella) in sand dunes, Namib Desert, Namibia
Grey heron (Ardea cinerea) catching fish in lake, Pusztaszer, Hungary, April
Killer whale (Orcinus orca) adult male surfacing at dusk surrounded by birds, who watch for herrings (Clupea harengus) left after they have fed, Andenes, Andoya island, North Atlantic Ocean, Norway
Praying mantis nymph silhouetted through leaf, . Raja Ampat, Western Papua, Indonesian New Guinea
Chinese lanterns (Physalis alkekengi) skeletons, silhouette on white background
Hedge parsley (Torilis japonica) close up pattern of dead seed heads. Silhouette on white background
Chinese Lanterns (Physalis alkekengi), Hydrangea flowers, Poplar leaves (Populus) and Honesty skeletons (Lunaria annua), silhouettes on white background
Ripe Barley (Hordeum vulgare) ears black and white backlit image
Stonehenge at sunrise, Salisbury Plain, Wiltshire, England, UK, June 2006
Corfe Castle in morning mist, Corfe, Isle of Purbeck, Dorset, England, UK. September 2013
Puffin (Fratercula arctica) wings spread backlit, Great Saltee Island, County Wexford, Republic of Ireland, June
Bottlenosed dolphin (Tursiops truncatus) porpoising during annual sardine run, Port St Johns, South Africa. June