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RF - Scarlet macaw (Ara macao) pair preening, Osa Peninsula, Costa Rica
Northern rockhopper penguin (Eudyptes moseleyi) portrait, Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Gentoo Penguin (Pygoscelis papua) coming out of the sea, Cuverville Island, Antarctic Peninsula, Antarctica
Barn owl (Tyto alba) flying through heavy snowfall, Derbyshire, February
Great egret (Ardea alba) flying across lake, Ranthambhore National Park, India
Green-winged macaws (Ara chloroptera) preening each other. Brazil. South America
Toco Toucan (Ramphastos toco) beak open with tongue visible while feeding on mango, Brazil
Atlantic Puffin (Fratercula arctica) in flight, coming into land with beak full of sand eels (Ammodytes tobianus). Fair Isle, Shetland Islands, Scotland, UK, July
Emperor penguin (Aptenodytes forsteri), chick in brood pouch, Snow Hill Island, Antarctic Peninsula
Arctic tern (Sterna paradisaea) in flight with fish in beak for feeding chick, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, July
Sword billed hummingbird (Ensifera ensifera) in flight, Guango, Napo, Ecuador
Rhinoceros hornbill (Buceros rhinoceros) male inflight, carrying mouse prey back to nest, Budo-Sungai Padi National Park, Thailand
Jackdaw (Corvus monedula) pair chasing a Grey squirrel (Sciurus carolinensis) as it emerges from a nest box the birds want to nest in which is already occupied by the squirrel and its mate
Jackdaw (Corvus monedula) hovering above a nest box it wants to nest in which a Grey squirrel (Sciurus carolinensis) and its mate have occupied, with the squirrel threatening the bird, Wiltshire, UK
Mute swan (Cygnus olor) moulting and plucking its own feathers, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. August
Baltimore oriole (Icterus galbula) female perched on Berlandiers fiddlewood (Citharexylum berlandieri). South Padre Island, Texas, USA. April
Long-tailed potoo (Nyctibius aethereus) camouflaged on roosting perch in lowland rainforest, Manu Biosphere Reserve, Peru
Great crested grebe (Podiceps cristatus) carrying chicks on its back in early morning light, Valkenhorst Nature Reserve, The Netherlands, Europe. May
Three Galah (Eolophus roseicapillus) drinking at inland waterhole, Cunnamulla, Queensland, Australia
Sparrowhawk (Accipiter nisus) juvenile flying, Parainen Uto, Finland. September
Goshawk (Accipiter gentilis) flying in snow, Mainua Kajaani, Finland. March
Waved albatross (Phoebastria irrorata) head portrait, Espanola Island, Galapagos, South America
Annas hummingbird (Calypte anna) male dancing in flight and displaying his bright pink throat feathers to a female, Montana, USA. June
Mute swan (Cygnus olor) swimming on sun-dappled water, Lower Silesia, Poland
European bee-eater (Merops apiaster) two perched on branch, Lower Silesia, Poland
Great white egret (Ardea alba) fishing in shallow pond, Valkenhorst Nature Reserve, The Netherlands, August
Mandarin duck (Aix galericulata) drake standing on frozen pond during snowstorm. Richmond Park, London, UK. January
Male Black woodpecker (Dryocopus martius) in snow, showing red crown. Kuusamo Finland December. Overall winner of the Swiss Vogelwarte photo competition 2021
Greater Prairie-Chicken (Tympanuchus cupido) male displaying on lek. Fort Pierre National Grassland, South Dakota, USA. When displaying
Common kingfisher (Alcedo atthis) perched on a bulrush, Devpm, UK, November
Great horned owls (Bubo virginianus), adult and chick, on nest in Saguaro cactus (Carnegiea gigantea), Sonoran desert, Arizona
Tawny owls (Strix aluco) peering out from tree hollow, Dumfries and Galloway, Scotland, UK, May, 2021
Tawny owl (Strix aluco) perched in Silver Birch tree (Betula pendula), Dumfries and Galloway, Scotland, UK, May
Superb lyrebird (Menura novaehollandiae) male in partial display perched in Tree fern (Cyatheales) in wet eucalypt forest, Lakes Entrance Rainforest, Victoria, Australia
Crested tit (Lophophanes cristatus), Sierra de Grazalema Natural Park, Southern Spain. September
Mandarin duck (Aix galericulata) male swimming with autumn colours reflected in the water, London, UK, November
Song thrush (Turdus philomelos) looking for worms in a garden, Broxwater, Cornwall, UK. April
Great horned owl (Bubo virginianus) adult and chick in nest in Saguaro cactus (Carnegiea gigantea) Sonoran Desert, Arizona, USA
Bearded vulture (Gypaetus barbatus) flying, Ordesa National Park, Pyrenees, Spain
Two Long-eared owl chick (Asio otus) perched in tree. Alsace. France
Baya weaver (Ploceus philippinus) breeding male perched on nest, rural Karantaka, India
An adult King penguin (Aptenodytes patagonicus) amongst a creche of chicks at the breeding colony. Fortuna Bay, South Georgia Island
A breeding pair of King penguins (Aptenodytes patagonicus) with their chick. St Andrews Bay, South Georgia Island
King penguins (Aptenodytes patagonicus) congregate on the beach, seeking a safe point to go into the sea, with male Southern elephant seals (Mirounga leonina) blocking their path
Yellow warbler (Setophaga petechia), male among red Sessuvium edmonstoni plants, San Cristobal Island, Galapagos Islands
Red-footed booby (Sula sula), brown morph, Genovesa Island, Galapagos Islands
Galapagos penguin (Spheniscus mendiculus), pair courting, Isabela Island, Galapagos Islands
Galapagos hawk (Buteo galapagoensis), juvenile plumage, Bainbridge Crater, Galapagos Islands
