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Lady Amhersts pheasant (Chrysolophus amherstiae) male displaying, Kanding, Sichuan, China. April
Bee-eater (Merops apiaster) male offering wasp as nuptial gift, Hungary. June
Common Cuckoo (Cuculus canorus) perched on gate Surrey, England, UK. April
Starling (Sturnus vulgaris) murmuration at sunset, West Pier, Brighton, England, UK
Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May
Cape buffalo (Syncerus caffer caffer) portrait, Red-billed oxpeckers (Buphagus erythrorhynchus) removing parasites. Nakuru, Kenya
Turtle dove (Streptopelia turtur) female feeding chicks on nest. Ranmore Common, near Dorking Surrey, England, UK. July
Canada Goose (Branta canadensis) in flight. Gilbert Water Ranch, Arizona, USA. January
Mandarin duck (Aix galericulata) pair mating on lake. Shanghai Zoo, China
Great Indian Hornbill (Buceros bicornis) in forest habitat, Tamil Nadu, Western Ghats, India
Grey francolin (Francolinus pondicerianus) Keoladeo Ghana National Park, Bharatpur, Rajasthan, India
Common blackbird (Turdus merula) feeding on Rowan (Sorbus aucuparia) berries, Leon, Spain, February
Bald eagle (Haliaeetus leucocephalus) portrait, captive, occurs in North America
Great tit (Parus major), perched on tree, Monmouthshire, Wales, UK. December
Rock Ptarmigan (Lagopus muta) portrait, Inari, Finland, September
Red-footed booby (Sula sula), adult and chick at nest. Genovesa Island, Galapagos
Nazca booby (Sula granti) preening. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Northern rockhopper penguin (Eudyptes moseleyi) parent with chick. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Andean condor (Vultur gryphus), Male, Antisanilla Reserve, Ecuador
Emperor penguin (Aptenodytes forsteri) tobogganing, Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November
Blue-footed booby (Sula nebouxii) landing, Santa Cruz Island, Galapagos, Ecuador
Stewart Island Brown Kiwi / Southern Tokoeka (Apteryx australis lawryi), female probing by scent for sand hoppers in rotting kelp beach wrack, Ocean Beach, Stewart Island, New Zealand
Okarito Brown Kiwi (Apteryx rowi) male known as Scooter patrolling his territory, from population of 200. Okarito Forest, Westland, South Island, New Zealand
Great Spotted Kiwi (Apteryx hstii) foraging in rainforest habitat at night, NW Nelson, South Island, Otorohanga Breeding Facility, New Zealand
Stellers jay (Cyanocitta stelleri) perched on branch. British Columbia, Canada, June
Chestnut-eared aracari (Pteroglossus castanotis) on cat-gut vine. Pantanal, Brazil, August
Common grackle pair (Quiscalus quiscula) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Osprey (Pandion haliaetus) with Atlantic needlefish (Strongylura marina) it has caught. Blue Cypress Lake, Florida, USA, April
Violet-tailed sylph hummingbird (Aglaiocercus coelestis) in flight, about to land. Ecuador, May
Small flock of Lapwings (Vanellus vanellus) flying in to rest after migration. Cresswell Pond, Druridge Bay, Northumberland, England, UK, September
Common buzzard (Buteo buteo) landing on prey, wings open, Scotland, UK, September 2016
Snares island crested penguin (Eudyptes robustus) colony on the coast, high angle view, Snares Island, New Zealand
Golen eagle (Aquila chrysaetos) adult with nest material on eyrie with chick showing background, Isle of Lewis, Scotland, UK. May
Puffins (Fratercula arctica) Shaint Isles, Scotland, UK, June
Grey heron (Ardea cinerea) hunting for food amongst moss with loch and Celtic rainforest in back ground, Scotland, UK, October
Mandarin duck (Aix galericulata) male female pair, Reddish Vale Country Park, Greater Manchester, UK. December
Emu (Dromaius novaehollandiae) juvenile, Cleland Wildlife Park, South Australia, March. Captive
Rainbow lorikeet (Trichoglossus moluccanus) on a branch, Cania Gorge National Park, Queensland, Australia. September
Green sandpiper (Tringa ochropus) with small worm it has caught in shallow freshwater lake, Gloucestershire, UK, August
Canada geese (Branta canadensis) in flight at Abbotsbury Swannery, Abbotsbury, Dorset, England, UK. August 2014
Three Little owls (Athene noctua) looking out of a nest hole, Cumbria, UK, August. Captive
Wood pigeons (Columba palumbus) male and female during courtship preening display, Norfolk, England, UK, August
Spotted flycatcher (Muscicapa striata) feeding a chick which has just left its nestbox with a solitary bee, Cornwall, UK, August
Fernandinas flicker (Colaptes fernandinae), Cuba, endemic species
Blue-eared kingfisher (Alcedo meninting) Sabah, Malaysia
Tufted duck (Aythya fuligula) male drake, Valkenhorst Nature reserve, Valkenswaard, The Netherlands, May
Great white egret (Egretta Alba) preening its feathers, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, July