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RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Red-and-green macaw (Ara chloropterus) flock feeding on wall of clay lick. Manu Biosphere Reserve, Amazonia, Peru
Blackbird (Turdus merula), male singing whilst perched in Ash (Fraxinus excelsior) tree, near Bath, England, UK. May
European roller (Coracias garrulus) peering out of nest with feather, Lake Csaj, Pusztaszer, Kiskunsagi National Park, Hungary, February
Northern raven (Corvus corax) pair perching on branch. Danube Delta, Romania, May
White tailed sea eagle (Haliaeetus albicilla) perched on tree stump with fish. Hooded crow (Corvus cornix) flying in to attack in background. Danube Delta, Romania. May
South Georgia pintail (Anas georgica georgica) swimming in front of Southern elephant seal (Mirounga leonina), male. Gold Harbour, South Georgia. October
Elephant (Loxodonta africana), male standing on hind legs to reach acacia pods with Cattle egrets (Bubulcus ibis) in background. Mana Pools National Park, Zimbabwe
Bee-eater (Merops apiaster) with Bumblebee prey, Sierra de Grazalema Natural Park, southern Spain, July
Iberian lynx (Lynx pardinus) with Magpie (Pica pica) behind, Sierra de Andujar Natural Park, Jaen, Spain, September
Great white egret (Ardea alba) group of three in winter, Kiskunsag National Park, Hungary. Winner of the Bird Category of Nature Photographer of the Year Awards 2018
White-tailed eagle (Haliaeetus albicilla) male and female perched on tree in snow, near Kuusamo, Finland. March. Runner up in the Birds category of the GDT European Wildlife Photographer of the Year
Victoria crowned pigeon (Goura victoria) head portrait, vulnerable species, captive
Shag (Phalacrocorax aristotelis) in flight over sea. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Razorbill (Alca torda) landing, Great Saltee Island, County Wexford, Ireland, June
Puffin (Fratercula arctica) wings spread backlit, Great Saltee Island, County Wexford, Republic of Ireland, June
Snowy owl (Bubo scandiacus) in flight, Ontario, Canada, January
Wild Buffons macaw x Scarlet macaw (Ara ambiguus x Ara macao) hybrid, Costa Rica
RF - Marabou (Leptoptilos crumenifer) with Whitebacked vultures (Gyps africanus) Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Red-billed blue magpie (Urocissa erythrorhyncha) perched on branch, Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Amazon kingfisher (Chloroceryle amazona) with fish, Cuiaba, Pantanal Matogrossense National Park, Pantanal, Brazil
California condor (Gymnogyps californianus). in wild, Baja, Mexico
Northern goshawk (Accipiter gentilis) perched on snag with grouse prey, Finland, March. March
Willow ptarmigan (Lagopus lagopus) two in flight, Taymyr Peninsula, Siberia, Russia. May
Western capercaillie (Tetrao urogallus) Tver, Russia. May
Black Grouse (Tetrao tetrix) displaying at a lek in snow, Tver, Russia. April
Rock ptarmigan (Lagopus muta) male on snow drift, Taymyr Peninsula, Siberia, Russia. March
Painted stork (Mycteria leucocephala), resting on nest with young at sunrise. Keoladeo / Bharatpur National Park UNESCO World Heritage Site, India
Chinese Sparrowhawk (Accipiter soloensis) flying Guangshui, Hubei province, China. July
Common hoopoe (Upupa epops) sitting on a stone, Tangjiahe National Nature Reserve, Sichuan Province, China
White-naped crane (Grus vipio) flying, Inner Mongolia, China
Lady Amhersts pheasant (Chrysolophus amherstiae) male walking on ground in Baihualing, Gaoligongshan, Yunnan, China
Magnificent hummingbird (Eugenes fulgens) female, Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Fiery throated hummingbird (Panterpe insignis) an endemic bird species. Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Mangrove hummingbird (Amazilia boucardi) female feeding chicks in nest, Pacific coast mangroves area, Costa Rica, Endangered species
Northern gannet (Morus bassanus) pair, courtship display, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, May
Semipalmated sandpiper (Calidris pusilla) flock in flight over water, Johnsons Mills Reserve, Bay of Fundy, New Brunswick, Canada, July
Reed warbler (Acrocephalus scirpaceus) with insect larvae in beak, Greylake RSPB Reserve, Somerset Levels, England, UK, June
American flamingo (Phoenicopterus ruber) pair in courtship, group of three feeding, Punta Cormorant, Floreana Island, Galapagos
Brown pelican (Pelecanus occidentalis) in flight, Turtle Cove, Santa Cruz Island, Galapagos
Waved albatross (Phoebastria irrorata) group of three on nest, Punta Suarez, Espanola Island, Galapagos
Galapagos penguin (Spheniscus mendiculus) incubating egg, Puerto Pajas, Isabela Island, Galapagos
Barn swallow (Hirundo rustica) feeding young, Monmouthshire, Wales, UK, July
Swallow-tailed gull (Creagrus furcatus) calling, Genovesa Island, Galapagos
Wood pigeon (Columba palumbus) pair preening one another, The Netherlands
Cliff parakeets (Myiopsitta luchsi) perched on cactus, Red-fronted Macaw Community Nature Reserve, Omerque, Bolivia
Mountain Caracara (Phalcoboenus megalopterus) taking off, Laguna Canapa, Altiplano, Bolivia
RF - Mute swan (Cygnus olor) taking off from a pond Valkenhorst Nature Reserve, Valkenswaard, the Netherlands June
