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English oak {Quercus robur} leaf in autumn, Donisthorpe, The National Forest, Leicestershire, UK. November
Red deer (Cervus elaphus) stag bellowing during autumn rut, Richmond Park, London, England, UK, October
Red deer (Cervus elaphus) young stag tasting the air (flehmen response) Bradgate Park, Leicestershire, England, UK, October
Birch trees blowing in high winds, long exposure, Calke Abbey, The National Forest, Central England, UK, November 2010
RF - Portrait of domestic German shorthaired pointer pup in autumn. Pomfret, Connecticut, USA. October
Birch trees in autumn, Kyle of Tongue, Sutherland, Scotland, UK, June 2017
Falls of Truim running through autumnal woodland, Cairngorms National Park, Scotland, UK, October 2016
Waterfall on wooded hillside in autumn, Glenfeshie, Cairngorms National Park, Scotland, UK, October 2015
Autumn colours and stormy clouds reflected in the calm waters of Loch Insh, Cairngorms National Park, Scotland, UK, November 2015
Quaking aspen trees (Populus tremuloides) and conifers with dusting of snow, Dome Hill above Dawson City, Yukon Territories, Canada, September 2013
Fallow deer (Dama dama) buck bellowing at dawn during the rut, Cheshire, UK, October
Close-up of group of toadstools (Mycena sp) autumn. Leicestershire, UK. November. Photographer quote: It is all too easy to overlook the fascinating miniture world underneath our feet - mosses
Grey wolf portrait with autumn leaves {Canis lupus} captive, USA
Grey wolf {Canis lupus} head profile, captive, Montana, USA
Spruce grouse (Falcipennis canadensis) in forest, Denali National Park, Alaska, USA, September
Eagle owl (Bubo bubo) in flight through forest, backlit at dawn, Czech Republic, November. Captive
Long-eared owl (Asio otus) perched in tree amongst autumn leaves, Czech Republic, October. Captive
English oak (Quercus robur) acorn. Cambridgeshire, UK. September
Red Squirrel (Sciurus vulgaris) leaping onto log, Scotland, UK, September. Highly commended in the Habitat category of the BWPA (British Wildlife Photography Awards) 2016
Red deer (Cervus elaphus) female amongst autumn foliage, Arran, Scotland, October 2013
Hedgehog (Erinaceus europaeus) amongst leaves in low sunlight. Leicestershire, UK, November
Red squirrel (Sciurus vulgaris) standing on its hind legs, Cairngorms National Park, Scotland, UK, March. Non-ex
Red deer (Cervus elaphus) hind and young calf, Bradgate Park, Leicestershire, England, UK, November
Red deer (Cervus elaphus) dominant stag amongst bracken, Bradgate Park, Leicestershire, England, UK, October. Did you know
Red Deer (Cervus elaphus) doe, silhouetted at dusk, Bradgate Park, UK, November
RF- Red squirrel in autumn (Sciurus vulgaris). Strathspey, Scotland, UK
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
Long-eared owl (Asio otus) in the rain, perched with feathers puffed up, on larch, captive, Yorkshire, UK
Raven (Corvus corax) calling in flight along cliff face, South Stack RSPB reserve, Anglesey, Wales, UK, October
RF- Chanterelle mushroom (Cantharellus cibarius), gill detail
American Bison (Bison bison) in habitat, Grand Teton National Park, Wyoming, USA, October
Great Tit (Parus major) perched on branch with yellow from autumn trees in the background. Southern Norway, October
Fallen leaves in pond with aquatic plants, Denmark, Europe, September
RF - Ripe black grapes Regent variety with autumn coloured leaves, Norfolk, England, UK, September
RF- Close up of Leaves of Red Oak (Quercus rubra) in autumn. October
Long-eared owl (Asio otus) in Rowan tree with red berries, captive, Yorkshire, UK
Dombai-Ulgen peak with Rhododendron luteum plant, Teberdinskii reserve, NW Caucasus, Russia
Hraunfossar waterfall in autumn, Iceland, September 2013
Ellis River high in White Mountain National Forest. New Hampshire, October 2012
Newlands Valley and Causey Pike, Lake District National Park, Cumbria, England, UK. October 2012
The Canal du Midi near Castelnaudary, Languedoc-Rousillon, France. September 2011
Quinling Golden snub nosed Monkey (Rhinopitecus roxellana qinligensis), infant sitting in a tree. Zhouzhi Nature Reserve, Qinling Mountains, Shaanxi, China
Lake Ann surrounded by Douglas firs (Pseudotsuga menziesii) Silver firs (Abies alba) and other conifers in the area of Rainy Pass of North Cascades National Park, North Cascades National Park
Fallow deer (Dama dama) in woodland clearing at dawn, Cheshire, UK November
Red deer (Cervus elaphus) stags fighting at dusk during rutting season, Cheshire, UK October
English oak tree (Quercus robur) fallen acorns and leaves on a bed of moss, Oxfordshire, UK October
Beech trees in autumn mist, Morvan regional park, Burgundy, France, November 2010
Virginia creeper (Parthenocissus quinquefolia), climbing on Live Oak (Quercus virginiana) with fall colors, Kerrville, Hill Country, Central Texas, USA, November