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Bramble leaves (Rubus plicatus) and Bracken fronds (Pteridium sp) changing colour in autumn, Norfolk, England, October
Beech woodland surrounding Burbage Brook. Padley Gorge, Peak District National Park, UK. Autumn, November 2010
Malham, Yorkshire Dales, England, UK. October 2011
Autumn colour in the trees blowing in the wind, The Promenade, Clifton Downs, Bristol, England, UK
Male Adder (Vipera berus) coiled on leaf litter. Cannock Chase, Staffordshire, UK, October
Quinling Golden snub nosed monkey (Rhinopitecus roxellana qinlingensis) family group foraging along a small creek in a gullly. Zhouzhi Nature Reserve, Qinling Mountains, Shaanxi, China
Landscape view of Vosges Forest at dawn, France, November 2011
Water running over rocks, Lodore Falls, with autumnal trees in the background, Lake District, Cumbria, England, UK, November
Oxbow Bend of the Snake River with the Grand Tetons on the horizon, Grand Teton National Park, Wyoming, USA, September 2006
Silver birch (Betula pendula) woodland in autumn, Glen Moriston, Scotland, UK, October 2011
Silver birch (Betula pendula) leaves in autumn, Glen Affric, Highland, Scotland, UK, October
Rainbow over Loch Beinn a Mheadhoin, Glen Affric, Highland, Scotland, UK, October 2010
Autumn leaves in a misty Beech (Fagus) woodland. Saint Gobain, France, November 2010
Scattered Scots pine trees (Pinus sylvestris) beside the Dorback Burn, Dorback Estate, Cairngorms National Park, Scotland, UK, September 2011
Red Fox (Vulpes vulpes) behind a tree and autumn leaves. Black Forest, Germany, November
Red deer (Cervus elaphus) three females or hinds in silhouette in dawn mist, Leicestershire, UK, October. Non-ex
Autumns colours in Carpathian beech forest. Bieszczady, Carpathian Mountains, Poland, November
Woodland stream flowing through Wilmersham Wood, Exmoor National Park, Somerset, UK. November 2010
Grey squirrel (Sciurus carolinensis) foraging in an oak woodland. Perthshire, Scotland, Nov
Autumn reflections on Modene Lake and forest. La Mauricie National Park, Quebec, Canada, October 2010
Red deer (Cervus elaphus) hind in early morning mist, Leicestershire, UK. October. (non-ex)
Pattern of green yellow, and red Maple (Acer sp. )leaves in autumn, North Chagrin ReservationPattern of green yellow, and red Maple (Acer sp.)leaves in autumn, North Chagrin Reservation, Ohio, USA, October
Group of Wild Boar (Sus Scrofa) trotting through forest of Rambouillet, near Paris, France. Autumn
Autumn colours of the Aspen trees (Populus tremula) and conifers in the snow, near Muleshoe, Bow Valley Parkway, Banff National Park, Alberta, Canada. October 2009
Golden Scalycap fungi (Pholiota aurivella) on Beech tree, Sussex, England
Goldfinch (Carduelis carduelis) perched on Hawthorn, Cheshire, UK, October
Silver birch trees {Betula pendula} in autumn, Glen Strathfarrar NNR, Scotland, UK, October
European beech (Fagus sylvatica) trees in autumn, Pollino National Park, Basilicata, Italy, November 2008
Male Japanese macaque (Macaca fuscata) roaring to attract females during the breeding season, Jigokudani, Joshinetsu Kogen NP, Nagano, Japan
Chestnuts from Sweet chestnut tree {Castenea sativa} Redes NP, Asturias, Northern Spain, October 2007
Grey squirrel {Sciurus carolinensis} adult feeding among the fallen leaves of Horse Chestnut tree, Derbyshire, UK
Lake and mixed woodland in autumn, Upper Peninsula, Michigan, USA
Upland Sugar Maple (Acer saccharum) forest in autumn, Porcupine Mountains State Park, Upper Peninsula, Michigan, USA
Autumnal trees in the Valley of Varrados, Val d Aran, Catalonia, Pyrenees
Beechwood in Ordesa y Monte Perdido National Park in the Pyrenees, Huesca, Spain
Waterfall in autumnal beechwood in Montseny Natural Park, Barcelona, Spain
Collared earth star (Geastrum triplex), two on woodland floor. South Yorkshire, England, UK. September
Mountain view towards Loch Nevis overlooking Knoydart peninsula with Skye beyond. Taken from Sgurr Cos na Breachd-laoidh, Lochaber, Scotland, UK, September 2016
Red deer (Cervus elaphus) male stag silhouetted at dusk, rut season, Jura, Scotland, UK, September
Bay cup fungus (Peziza badia) Peak District National Park, Derbyshire, UK September
Weasel (Mustela nivalis) investigating birch stump with bracket fungus in autumn woodland, Sheffield
Sulphur Tuft Fungus (Hypholoma fasciculare) growing in clusters, Sussex, England, UK, September
Three 10 month old Badgers {Meles meles} browsing in leaf litter at night, Derbyshire, UK
Giant Polypore fungus (Meripilus giganteus) growing in clusters on woodland floor, UK, August
Ripening blackberries on Bramble bush (Rubus plicatus) Somerset, UK
Wirehaired daschund, Staffordshire bull terrier and Hungarian viszla in autumn colours, Wiltshire, UK
Brentor Church, Dartmoor, Devon. October 2009
Blue jay (Cyanocitta cristata) holding an acorn in its bill whilst perched on tree stump, with autumn foliage, New York, USA, October