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Coastal agave (Agave shawii) leaves. Near Bahia de Los Angeles, Baja California, Mexico
Lord Howe Island stick insect (Dryococelus australis), Bristol Zoo Gardens, Bristol, UK
Lemon shark pups (Negaprion brevirostris) in mangrove forest, Eleuthera, Bahamas
Flock of ducks silhouetted against sunset flying over field in winter in the Uitkerkse Polder nature reserve near Blankenberge, West Flanders, Belgium. February 2020
Species rich alpine meadow with Orange lily (Lilium bulbiferum), Meadow clary (Salvia pratensis) and Yellow rattle (Rhinathus sp)
North American Beaver (Castor canadensis) grooming. Acadia National Park, Maine, USA
Corn snake (Pantherophis guttatus), female with recently laid eggs, captive, native to Eastern United States
Keel-billed toucan (Ramphastos sulfuratus) feeding, tossing fruit seed in beak, perched on branch. Boca Tapada, Costa Rica. Sequence 2/2
Yellow bellied snapping turtle (Elseya flaviventralis) about to surface for air, Arnhem Escarpment Creek, Northern Territory, Australia. June
Apple hawk-moth (Langia zenzeroides formosana) Nzaus Mamei Jianshih Village, Hsssinchu County, Taiwan, Controlled conditions
Pine marten (Martes martes) standing on hind legs, Vosges, France
Coastal agave (Agave shawii) flowering in Sonoran Desert. Near Bahia de Los Angeles, Baja California, Mexico
Boojum tree (Fouquieria columnaris) and Mexican giant cardon (Pachycereus pringlei) amongst rocks in Sonoran Desert, at sunset. Baja California, Mexico. 2017
California condor (Gymnogyps californianus) landing, forest in background. Bird with tags on wings. Near San Pedro Martir National Park, Northern Baja California, Mexico. 2017
Mexican giant cardon cactus (Pachycereus pringlei) and Boojum tree (Fouquieria columnaris) in Sonoran Desert, rainbow in background. Catavina, Valle de los Cirios Reserve, Baja California, Mexico
Common collared lizard (Crotaphytus collaris auriceps) male basking on rock. Arches National Park, Utah, USA. May
Snow covered Mount Chephren, coniferous forest below. Banff National Park, Alberta, Canada. October 2018
Spotted eagle ray (Aetobatus narinari), Kicker Rock, San Cristobal Island, Galapagos
Walrus (Odobenus rosmarus) amongst group hauled out on shore. Spitsbergen, Svalbard, Norway. July
Aboriginal rock art, Wandjina style, Raft Point, Kimberley, Western Australia
Abandoned red mud (alumina) pond, where highly alkaline waste product produced by the industrial production of aluminium was stored
Worrells turtle (Emydura subglobosa worrelli), large adult female resting at night, Lawn Hill Creek, Queensland, Australia. January
Sandstone long-necked turtle (Chelodina burrungandjii) actively foraging at night, Pentecost River, Western Australia. April
Close up of a Lord Howe Island stick insect (Dryococelus australis), Bristol Zoo Gardens, Bristol, UK. Captive
Bedstraw hawk-moth (Hyles gallii) Wallis, Switzerland, April. Controlled conditions
Red Bird-of-Paradise (Paradisaea rubra) male performing practice display at tree-top lek, Raja Ampat, Indonesia. *Medium repro only
Atlantic puffin (Fratercula arctica) with beakful of fish, Saltee Islands, County Wexford, Republic of Ireland
View from Brandon Mountain Dingle, County Kerry, Ireland, July 2008
Scarce Merveille du Jour moth (Moma alpium) camouflaged against lichen, Beetzseeheide, Germany, March
Pico de Mondarruego and Arazas River Valle de Ordesa, Ordesa National Park, Spain, October
False tiger moth (Dysphania militaris) Isle of Marinduque, Philipines
Bumblebee (Bombus) on Rampion flower (Phyteuma), France, May
Common crane (Grus grus) flock feeding in field in snow, Champagne, France, February
Red squirrel (Sciurus vulgaris) climbing tree trunk in forest, Vosges, France, May
Roe deer (Capreolus capreolus) fawn, Vosges, France, June
Roe deer (Capreolus capreolus) doe, leaping whilst running away, rear view, Vosges, France
Badgers (Meles meles) at sett entrance, interacting, Vosges, France, May
Wild cat (Felis silvestris) Vosges, France
Badger (Meles meles) two outside sett interacting, Vosges, France, May
Badger (Meles meles) outside sett, play fighting, Vosges, France, May
Red deer (Cervus elaphus) doe in winter snow, Vosges, France, January
Crested drongo, (Dicrurus forficatus), Berenty National Park, Madagascar
Coastal agave (Agave shawii) leaves dying after flower has bloomed. North Baja California, Mexico
Stellers sea eagle, (Haliaeetus pelagicus), on sea pack ice, Okhotsk Sea, Rausu, Hokkaido, Japan
Coastal agave (Agave shawii), two flower spikes against sky, view from below. Near Bahia de los Angeles. Baja California, Mexico
Sour pitaya (Stenocereus gummosus) with Aloe (Aloe sp) flower spikes in background. Sonoran Desert, Puerto Balandra, near La Paz, Baja California Sur, Mexico. 2013
Mexican giant cardon (Pachycereus pringlei), two young cacti amongst flowering Desert sand verbena (Abronia villosa) during spring super bloom. Northern Baja California, Mexico
Desert lily (Hesperocallis undulata). Lower Colorado Desert, Northern Baja, Mexico
Hollowleaf annual lupine (Lupinus succulentus) flowering in front of rock. Catavina, Valle de los Cirios Reserve, Baja California, Mexico
Coastal century (Agave shawii), Baja elephant tree (Pachycormus discolor) and Boojum tree (Fouquieria columnaris) in Sonoran Desert. Near Bahia de Los Angeles, Baja California, Mexico
Hickman Bridge, rock arch in Navajo sandstone. Capitol Reef National Park, Utah, USA. February 2020
Orange lily (Lilium bulbiferum) and Meadow clary (Salvia pratensis) flowers in alpine meadow. Dolomites, Italy. June
Rhaetian poppy (Papaver rhaeticum) above lake in Dolomites at 2200m. Valparola Pass, near Cortina, Belluno, Italy. July 2016
Sulphur pasqueflower (Pulsatilla alpina apiifolia) in alpine grassalnd in Dolomites, mountain in background. Trentino, Italy. June 2019
Oxlip (Primula elatior) in alpine pasture following snow melt, mountains in background. Col di Rodella, Fassa Valley, Dolomites, Trentino, Italy. June 2019
The lighthouse La Vieille in the strait Raz de Sein at dusk at the Pointe du Raz, Plogoff, Finistere, Brittany, France. September 2019
Least Chipmunk (Tamias minimus) feeding on seeds, Grand Teton National Park, Wyoming, USA
Spotted eagle rays (Aetobatus narinari), in mangrove inlet, Elizabeth Bay, Isabela Island, Galapagos
Marbled Stingray (Taeniurops meyeni), Galapagos
Ogoy Island, Maloye More Strait. Lake Baikal, Siberia, Russia, April
Black-tailed godwit (Limosa limosa) flock in flight past Willow trees fringing a shallow lake, Gloucestershire, UK, February
Two wild konik horse stallions fighting in front of herd, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
A band of wild konik horses walking with newborn foal, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
A wild konik horse mare sniffing her yearling foal as he sniffs her newborn foal, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
Two wild Konik horse stallions greeting one another, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
A herd of wild konik horses running, with refection in water, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
Mountain primrose (Primula villosa) in rock crevice at 2300m in elevation, mountains in background. North Tyrol, Austrian. June
Highland cows, three silhouetted at dusk. Glengorm Estate, Isle of Mull, Scotland, UK. June
Golden-browed chlorophonia (Chlorophonia callophrys), two males perched on moss covered branch. Savegre Valley. Costa Rica
Silk moth (Copaxa sp) camouflaged against leaf litter on rainforest floor. Bosque de Paz, Costa Rica
Shining honeycreeper (Cyanerpes lucidus) male and Red-legged honeycreeper (Cyanerpes cyaneus) perched on branch with epiphyte. Boca Tapada, Costa Rica
Black-bellied bustard (Lissotis melanogaster) male calling at sunrise. Serengeti National Park, Tanzania
Conifers on peaks of Huangshan Mountains, in fog. Anhui Province, China. 2016
Yak (Bos grunniens) herd on plain beside river, mountains in background. Near Litang, Kham, Garze Tibetan Autonomous Prefecture, Sichuan, China. October 2016
Coniferous forest on mountainside in fog. Kham Mountains, Tibet. October 2016
Clouds in evening light. Kham, Tibet
Yak (Bos grunniens) grazing beside wall of Ganden Thubchen Choekhorling Monastery, mountains in background. Litang, Garze Tibetan Autonomous Prefecture, Sichuan, China. October 2016
Northern white-cheeked gibbon (Nomascus leucogenys), adult male, Xishuangbanna National Park, Yunnan, China. Critically endangered
Short eared rock wallaby with offspring (Petrogale brachyotis brachotis), Ord River, Kimberley, Western Australia
King George River iron stained sandstone cliffs, Kimberley, Western Australia
Turtle dove (Streptopelia turtur) pair, Teide National Park, Tenerife, Canary Islands, Spain, July
Tail of Blue whale diving (Balaenoptera musculus brevicauda) possibly the pygmy sub-species of blue whale, (Balaenoptera musculus). Mirissa, Sri Lanka, Indian Ocean. Photo taken under permit
Tail of Blue whale (Balaenoptera musculus brevicauda). This may be the pygmy sub-species of blue whale, Balaenoptera musculus. Mirissa, Sri Lanka, Indian Ocean
Forest with lush vegetation Portland, Oregon, USA. April