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Hickman Bridge, rock arch in Navajo sandstone. Capitol Reef National Park, Utah, USA. February 2020
Mexican giant cardon cactus (Pachycereus pringlei) and Boojum tree (Fouquieria columnaris) in Sonoran Desert, rainbow in background. Catavina, Valle de los Cirios Reserve, Baja California, Mexico
Orange lily (Lilium bulbiferum) and Meadow clary (Salvia pratensis) flowers in alpine meadow. Dolomites, Italy. June
Rhaetian poppy (Papaver rhaeticum) above lake in Dolomites at 2200m. Valparola Pass, near Cortina, Belluno, Italy. July 2016
Sulphur pasqueflower (Pulsatilla alpina apiifolia) in alpine grassalnd in Dolomites, mountain in background. Trentino, Italy. June 2019
Oxlip (Primula elatior) in alpine pasture following snow melt, mountains in background. Col di Rodella, Fassa Valley, Dolomites, Trentino, Italy. June 2019
The lighthouse La Vieille in the strait Raz de Sein at dusk at the Pointe du Raz, Plogoff, Finistere, Brittany, France. September 2019
Least Chipmunk (Tamias minimus) feeding on seeds, Grand Teton National Park, Wyoming, USA
Spotted eagle ray (Aetobatus narinari), Kicker Rock, San Cristobal Island, Galapagos
Spotted eagle rays (Aetobatus narinari), in mangrove inlet, Elizabeth Bay, Isabela Island, Galapagos
Marbled Stingray (Taeniurops meyeni), Galapagos
Ogoy Island, Maloye More Strait. Lake Baikal, Siberia, Russia, April
Black-tailed godwit (Limosa limosa) flock in flight past Willow trees fringing a shallow lake, Gloucestershire, UK, February
Two wild konik horse stallions fighting in front of herd, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
A band of wild konik horses walking with newborn foal, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
A wild konik horse mare sniffing her yearling foal as he sniffs her newborn foal, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
Two wild Konik horse stallions greeting one another, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
A herd of wild konik horses running, with refection in water, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
Mountain primrose (Primula villosa) in rock crevice at 2300m in elevation, mountains in background. North Tyrol, Austrian. June
Highland cows, three silhouetted at dusk. Glengorm Estate, Isle of Mull, Scotland, UK. June
Golden-browed chlorophonia (Chlorophonia callophrys), two males perched on moss covered branch. Savegre Valley. Costa Rica
Silk moth (Copaxa sp) camouflaged against leaf litter on rainforest floor. Bosque de Paz, Costa Rica
Shining honeycreeper (Cyanerpes lucidus) male and Red-legged honeycreeper (Cyanerpes cyaneus) perched on branch with epiphyte. Boca Tapada, Costa Rica
Black-bellied bustard (Lissotis melanogaster) male calling at sunrise. Serengeti National Park, Tanzania
Conifers on peaks of Huangshan Mountains, in fog. Anhui Province, China. 2016
Yak (Bos grunniens) herd on plain beside river, mountains in background. Near Litang, Kham, Garze Tibetan Autonomous Prefecture, Sichuan, China. October 2016
Coniferous forest on mountainside in fog. Kham Mountains, Tibet. October 2016
Clouds in evening light. Kham, Tibet
Yak (Bos grunniens) grazing beside wall of Ganden Thubchen Choekhorling Monastery, mountains in background. Litang, Garze Tibetan Autonomous Prefecture, Sichuan, China. October 2016
Northern white-cheeked gibbon (Nomascus leucogenys), adult male, Xishuangbanna National Park, Yunnan, China. Critically endangered
Short eared rock wallaby with offspring (Petrogale brachyotis brachotis), Ord River, Kimberley, Western Australia
King George River iron stained sandstone cliffs, Kimberley, Western Australia
Turtle dove (Streptopelia turtur) pair, Teide National Park, Tenerife, Canary Islands, Spain, July
Tail of Blue whale diving (Balaenoptera musculus brevicauda) possibly the pygmy sub-species of blue whale, (Balaenoptera musculus). Mirissa, Sri Lanka, Indian Ocean. Photo taken under permit
Tail of Blue whale (Balaenoptera musculus brevicauda). This may be the pygmy sub-species of blue whale, Balaenoptera musculus. Mirissa, Sri Lanka, Indian Ocean
Forest with lush vegetation Portland, Oregon, USA. April
Coastal wetlands of RSPB Titchwell Nature Reserve, Norfolk, UK, March
African fish eagle (Haliaeetus vocifer) attacking a female Saddle-billed Stork (Ephippiorhyncus senegalensis )trying to steal a fish it has just caught. Liuwa Plains National Park, Zambia
Snow leopard (Panthera uncia) female in snow, Hemis National Park, Ladakh, India
King Penguins (Aptenodytes patagonicus) approached by an Antarctic Fur Seal (Arctocephalus gazella). A few pecks from the penguins caused the seal to retreat
Lord Howe Island stick insect (Dryococelus australis) eggs developing in damp vermiculite, part of a breeding program at Bristol Zoo Gardens, Bristol, UK
Apple hawk-moth (Langia zenzeroides formosana), Hsssinchu County, Taiwan, Controlled conditions
Boojum tree (Fouquieria columnaris) in Sonoran Desert. Near Bahia de Los Angeles, Baja California, Mexico
California condor (Gymnogyps californianus) group in flight, tags and radio transmitters on wings. Baja, Mexico. 2017
California condor (Gymnogyps californianus) in flight. Near San Pedro Martir National Park, Northern Baja California, Mexico
Spotted eagle rays (Aetobatus narinari) silhouetted, Kicker Rock, San Cristobal Island, Galapagos
Keel-billed toucan (Ramphastos sulfuratus) looking downwards, perched on branch. Boca Tapada, Costa Rica