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Pair of Spotted unicornfish (Naso brevirostris) courtship display above the reef, Ari Atoll, Maldives, Indian Ocean
Pair of Nudibranchs (Flabellina exoptata) just prior to mating, the mating apparatus of the one on left is visible, Bitung, North Sulawesi, Indonesia, Molucca Sea
Two Ruffs (Philomachus pugnax) male, competing at a lek, Vardo, Norway. May
Pair of Couch's spadefoot toads (Scaphiopus couchii) mating in water, Texas, USA. June
Vermilion flycatcher (Pyrocephalus obscurus) male, courtship display, Texas, USA. May
American flamingo (Phoenicopterus ruber) performing courtship wing flashing, Quinta Playa, Isabela Island, Galapagos Islands, Ecuador
Magnificent bird of paradise (Cicinnurus magnificus) male, leaning back and presenting his breast shield and raised cape to two females, courtship display, Arfak Mountains, West Papua, Indonesia
Wilson's bird of paradise (Cicinnurus respublica) male, courtship display, Batanta Island, West Papua, Indonesia
Pair of Wilson's birds of paradise (Cicinnurus respublica), male performing courtship display to female from a small sapling in the middle of his display court, Batanta Island, West Papua
Vogelkop bowerbird (Amblyornis inornatus) male, at his bower surrounded by orange leaves and petals and other objects he has collected, Arfak Mountains, Vogelkop Peninsula, West Papua
Wandering albatross (Diomedea exulans) male, standing with wings spread in courtship display, Prion Island, South Georgia
Magnificent riflebird (Ptiloris magnificus alberti) male, courtship display, Piccaninny Plains Sanctuary, Cape York Peninsula, Queensland, Australia
Pair of Lava gulls (Leucophaeus fuliginosus) during courtship, Genovesa Island, Galapagos Islands, Ecuador
Flock of American flamingos (Phoenicopterus ruber) performing courtship strutting and wing flashing, Quinta Playa, Isabela Island, Galapagos Islands, Ecuador
Male Beautiful sunbird (Cinnyris Pulchellus) courtship display, Allahein River, The Gambia
Female Olive ridley turtle (Lepidochelys olivacea) coming ashore at dusk to lay eggs, Oaxaca, Mexico, Pacific Ocean
Female Olive ridley turtles (Lepidochelys olivacea) coming ashore at night in large numbers to lay eggs during arribada mass nesting, Oaxaca, Mexico
Pair of Peppermint basslets (Liopropoma rubre) spawning at sunset, with eggs floating alongside the female, Grand Cayman, Cayman Islands, Caribbean Sea
Two male Black grouse (Tetrao tetrix) fighting for territory on lek, Cairngorms National Park, Scotland, UK. May
Male Black grouse (Tetrao tetrix) performing courtship display at lek, Cairngorms National Park, Scotland, UK. April
Two male Black grouse (Lyrurus tetrix) fighting and sparing over territory at a snow covered lek, Vasterbotten, Sweden. April
Male Western capercaillie (Tetrao urogallus) calling during lek mating, Finland. March
Pair of adult Red-necked grebe (Podiceps grisegena) calling, Finland. May
Close up of male Western capercaillie (Tetrao urogallus) during lek mating, Finland. March
Jackdaw (Corvus monedula) pair dive bombing and harassing a Grey squirrel (Sciurus carolinensis) as it emerges from a nest box the birds want to nest in which is already occupied by the squirrel
Wolf spider (Pardosa sp. ) carrying spiderlings on her back. UKWolf spider (Pardosa sp.) carrying spiderlings on her back. UK
Pair of Great crested grebes (Podiceps cristatus) performing the weed dance during courtship, The Netherlands, Europe. March
Pair of Mandarinfish (Synchiropus splendidus) mating, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Female Yellow-footed rock wallaby (Petrogale xanthopus) fending off advances from male (right), Idalia National Park, Queensland, Australia
Red-throated diver (Gavia stellata) pair displaying on the water, Vaala, Finland. May
Pair of Waved albatross (Phoebastria irrorata) courtship, Espanola Island, Galapagos, South America
Silvery grebe (Podiceps occipitalis) attempting to join Hooded grebes (Podiceps gallardoi) during a breeding display, Patagonia National Park, Santa Cruz, Argentina. Cropped
Pair of Leopard tortoises (Stigmochelys pardalis) mating, Karoo, South Africa
White stork (Ciconia ciconia) pair displaying on their nest built on the chimneys of Knepp Castle, Sussex, UK, June
Northern gannets (Morus bassanus) pair displaying amongst campion, Bempton Cliffs, Yorkshire, UK, May
Pair of mating Big blue octopuses (Octopus cyanea) where the male, on the right, is fertilising the female using its fertilising tentacle. Indian ocean, Maldives
Gentoo penguins (Pygoscelis papua) pair, Sea Lion Island, Falkland Islands
Black-browed albatross (Thalassarche melanophris) pair in courtship. Falkland Islands
Waved albatross (Phoebastria irrorata), pair mutual preening in courtship, Espanola Island, Galapagos Islands
Kingfisher (Alcedo atthis) male passing fish to female, courtship behaviour, Lorraine, France, March
Pink salmon (Oncorhynchus gorbuscha) migrate up river, Vancouver Island, Canada
Bilingual frog (Crinia bilingua) adult male calling, Adelaide River, Northern Territory, Australia, January
Stag beetle (Lucanus cervus) males fighting at dusk in oak wood, the Netherlands, June
Two Black-browed albatross (Thalassarche melanophris) rub their bills together - part of a bonding ritual. West Point Island, The Falkland Islands
Aerial view of Lesser flamingos (Phoeniconaias minor), mating display Lake Magadi, Rift Valley, Kenya
Arctic tern (Sterna paradisaea) pair mating, on rocks on beach. Hailuoto, Finland. May
Red deer (Cervus elaphus) stags fighting during the rutting season, Bushy Park, London, UK October
White-lipped deer (Cervus albirostris) walking on the Tibetan plateau, Serxu, Garze Prefecture, Sichuan Province, China
