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Black grouse (Lyrurus tetrix), male at lek, Finland, April
Satin bowerbird male at bower decorated with blue objects to attract mate, Lamington NP
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Western capercaillie (Tetrao urogallus) Tver, Russia. May
Black Grouse (Tetrao tetrix) displaying at a lek in snow, Tver, Russia. April
Northern gannet (Morus bassanus) pair, courtship display, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, May
Wood pigeon (Columba palumbus) pair preening one another, The Netherlands
European praying mantis female eats male after mating. Spain {Mantis religiosa} Sequence 2 of 3
Capercailie (Tetrao urogallus) male displaying from branch of Scots pine tree. Cairngorms National Park, Highlands, Scotland, UK, March
Common goldeneye (Bucephala clangula) male performing its mating display. Southern Norway. March
Raggiana bird of paradise (Paradisaea raggiana) male displaying to female, Papua New Guinea, Varirata NP
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Mandarinfish (Synchiropus splendidus) pair spawning. The larger male (behind) is lifting the female into the spawning rise on his pectoral fin, the smaller female (foreground) is releasing eggs
Waved Albatross (Phoebastria irrorata) courtship, Punta Cevallos, Espaola Island, Galapagos Islands, Ecuador, South America. Endemic, Critically endangered
Green sea turtle female (Chelonia mydas) returning to the sea, Tortuguero National Park, Costa Rica
Green-winged macaws (Ara chloroptera) preening each other. Brazil. South America
Humpback whale (Megaptera novaeangliae australis) courtship, with the female above showing her abdomen and the male below. Vava'u, Tonga. Pacific Ocean
Kori bustard (Ardeotis kori) male, performing mating display, silhouetted at dusk, El Karama Wilderness Conservancy, northern Kenya
Two Great egrets (Ardea alba) standing opposite each other with Grey heron (Ardea cinerea) in between. Lake Csaj, Pusztaszer, Hungary, January
Male Black grouse (Tetrao / Lyrurus tetrix) displaying at lek. Kuusamo, Finland, April
Black Grouse (Tetrao tetrix) males at lek, Utajarvi, Finland, April
Jackdaw (Corvus monedula) pair chasing a Grey squirrel (Sciurus carolinensis) as it emerges from a nest box the birds want to nest in which is already occupied by the squirrel and its mate
Jackdaw (Corvus monedula) hovering above a nest box it wants to nest in which a Grey squirrel (Sciurus carolinensis) and its mate have occupied, with the squirrel threatening the bird, Wiltshire, UK
Annas hummingbird (Calypte anna) male dancing in flight and displaying his bright pink throat feathers to a female, Montana, USA. June
Firefly (Lamprohiza splendidula) light trails of males in forest at dusk, Bavaria, Germany. July
Greater Prairie-Chicken (Tympanuchus cupido) male displaying on lek. Fort Pierre National Grassland, South Dakota, USA. When displaying
Tench (Tinca tinca) during spawning period amongst European watermilfoil (Myriophyllum spicatum), Sion, Valais canton, southwestern Switzerland
Galapagos penguin (Spheniscus mendiculus), pair courting, Isabela Island, Galapagos Islands
Red fox (Vulpes vulpes) pair nuzzling. London, UK. February
Male Black grouse (Tetrao / Lyrurus tetrix) in flight. Kuusamo, Finland, April
Boat-tailed grackle pair (Quiscalus major) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Apollo butterfly (Parnassius apollo) female laying eggs on branch. Parainen, Finland. July
Lawess parotia (Parotia lawesii) male displaying. Papua New Guinea
Red Bird-of-Paradise (Paradisaea rubra) male performing practice display at tree-top lek, Raja Ampat, Indonesia. *Medium repro only
Kottigehar dancing frog (Micrixalus kottigeharensis), male waving foot and calling, territorial behaviour. Agumbe, Western Ghats, India. Endemic
Red Deer (Cervus elaphus) stags fighting during the rutting season. Bushy Park, London, UK. October
Giant barrel sponge (Xestospongia testudinaria) releases a large cloud of gametes as it spawns on a coral reef, while a Threadfin hawkfish (Cirrhitichthys aprinus) looks
Sockeye salmon (Oncorhynchus nerka) males fighting in river Adams River, British Columbia, Canada. October
Espanola saddelback tortoise (Chelonoidis hoodensis) pair mating, Espanola Island, Galapagos
Colorado river toad (Incilius alvarius) pair mating. Sonoran desert, Arizona, USA. August
Yellowtail tang (Zebrasoma xanthurum) male and female turn belly to belly during spawning to ensure fertilization. Temple, Ras Umm Sid, Sharm El Sheikh, Sinai, Egypt. Red Sea
Manchurian crane (Grus japonensis) pair in courtship dance during snowstorm. Hokkaido, Japan. March
Fireflies (Luciola parvula himebotaru) flashing at night for courtship and reproduction. Gifu, Japan. Composite image
Grey seal (Halichoerus grypus), male and female, mating behaviour, Heligoland, Germany
