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Little owls courtship {Athene noctua} Spain
Blue-footed booby (Sula nebouxii) courting pair, South coast, Santa Cruz Island, Galapagos
Blue footed booby displaying {Sula nebouxii} Galapagos
Male Leach's sea star (Leiaster leachi) broadcast spawning, releasing streams of sperm from its arms. Kagoshima Prefecture, Kyushu, Japan
Indo-Pacific bottlenose dolphin (Tursiops aduncus) with penis extended. Ogasawara / Bonin Islands, Japan
Japanese crane (Grus japonensis) displaying, wings in heart shape, Hokkiado, Japan, February
Australasian crested grebe pair (Podiceps cristatus australis) in courtship display. Ashburton Lakes, Canterbury, New Zealand. August
Great crested grebe (Podiceps cristatus) performing their weed dance during courtship. Den Oever, The Netherlands. July
Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
Vogelkop superb bird-of-paradise (Lophorina niedda) male, performing courtship display on his display log, flashing false blue eye spots, a bright yellow gape, and blue breast shield
Andean cock-of-the-rock (Rupicola peruvianus), male displaying at lek in tree. Manu Biosphere Reserve, Amazonia, Peru
Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December
Brown hare, (Lepus europaeus), group of animals in field, Islay, Scotland, UK. March
Mating pair of chalkhill blue butterflies (Lysandra coridon) with wings closed resting on Devils-bit scabious (Succisa pratensis), Hatch Hill, Somerset, UK. August
Bald eagle (Haliaeetus leucocephalus) pair flying with claws linked during courtship flight, Homer, Alaska, USA, March
Rainbow lorikeets (Trichoglossus moluccanus) pair preening each other, perched in tree. Ulladulla, New South Wales, Australia
Great crested grebe (Podiceps cristatus) pair performing courtship displaying at dawn, backlit and surrounded by mist, Cheshire, UK, April
Red deer stag (Cervus elaphus) calling during rut. Richmond Park, England, UK, Europe
Mother of thousands (Kalanchoe daigremontiana), viviparous growth with vegetative propagation of plantlets on leaf margins. In cultivation
Reticulated glass frogs (Hyalinobatrachium valerioi) pair in amplexus, Osa Peninsula, Costa Rica
Ultramel Okeetee corn snake, with recently laid eggs, an interspecies hybrid between a Corn snake, (Pantherophis guttatus), and a Grey rat snake (Pantherophis spiloides)
Timber rattlesnake (Crotalus horridus) female with newborn young, Pennsylvania, USA. September
Elk (Cervus canadensis) bull and cw sniffing noses, Yellowstone National Park, Wyoming, USA. September
Rocky mountain elk (Cervus elaphus nelsoni) bull bugling during rut. Rocky Mountain National Park, Colorado, USA. October
Red Bird-of-Paradise (Paradisaea rubra) male performing practice display at tree-top lek, Raja Ampat, Indonesia. *Medium repro only
African lion (Panthera leo) and lioness after mating, Masai Mara National Reserve, Kenya
Lady Amhersts pheasant (Chrysolophus amherstiae) male displaying, Kanding, Sichuan, China. April
Bee-eater (Merops apiaster) male offering wasp as nuptial gift, Hungary. June
Bush cricket / Katydid (Tettigoniidae ), female leaf mimic ovipositing into branch. Manu Biosphere Reserve, Peru
Fireflies (Luciola parvula himebotaru) flashing at night for courtship and reproduction. Gifu, Japan. Composite image
Humpback whales (Megaptera novaeangliae) engaged in courtship, with the male hovering above the female. Vava'u, Tonga, Pacific Ocean
Twinspot snapper (Lutjanus bohar) shoal rushing up to spawn, releasing a cloud of milky white sperm and eggs in the water column. Caroline Islands, Palau, November
Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May
Mandarin duck (Aix galericulata) pair mating on lake. Shanghai Zoo, China
Green turtle (Chelonia mydas) female laying eggs, North coast, Santa Cruz Island, Galapagos
Oilbird (Steatornis caripensis) nocturnal birds roosting and nesting in along walls of deep, narrow gorge in farmland, Tandayapa Valley, 1500m altitude, Ecuador
Common grackle pair (Quiscalus quiscula) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Sarus crane (Grus antigone), male displaying, Keoladeo NP, Bharatpur, India
Wood pigeons (Columba palumbus) male and female during courtship preening display, Norfolk, England, UK, August
Mandarin duck drake (Aix galericulata) on the water, showing orange sail feathers which are used when displaying, Southwest London, UK, March
Queen conch (Lobatus gigas) laying eggs in the Exuma Cays Land and Sea Park, Exuma, Bahamas
Clarks grebes (Aechmophorus clarkii) courting pair performing the Weed Ceremony in which they slowly circle breast to breast holding pond weed as nesting material, Lake Hodges, Escondido, California
Black-necked stilts (Himantopus mexicanus) pair performing post-copulatory display (as copulation ends, the male slides off the female leaving his wings draped over her)
Black terns (Chlidonias niger), pair on a perch, vocalizing during courtship (termed Erect Posture display), Montezuma National Wildlife Refuge, New York, USA, May
White stork (Ciconia ciconia) pair mating on their nest at sunset, Knepp estate, Sussex, UK, April 2019. This is the first recorded instance of White storks nesting in the UK for several hundreds of
Red-footed booby (Sula sula), pair perched in tree. Genovesa Island, Galapagos
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Great bowerbird (Chlamydera nuchalis), male showing off treasures whilst female inspects bower. Lake Argyle, Kununurra, Western Australia