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Waved albatross (Phoebastria irrorata) pair in courtship display at nest site, Punta Suarez, Espanola Island, Galapagos
Black-browed albatross pair (Thalassarche melanophris) mutual grooming during copulation. New Island, Falkland Islands, November
Nurse sharks (Ginglymostoma cirratum) three in a courtship dance at sunrise in a mangrove area near Eleuthera, Bahamas
Magellanic penguin (Spheniscus magellanicus) calling at the entrance of burrow, courtship display, Carcass Island, Falkland Islands, October
Laysan albatross (Phoebastria immutabilis), courtship ritual, Eastern island, Midway Atoll National Wildlife Refuge, Hawaii
Common toad (Bufo bufo) pair in amplexus, Deeside, Scotland April
Iolas blue butterfly (Iolana iolas) pair mating on flowers, Aosta Valley, Gran Paradiso National Park, Italy
Brown Hare (Lepus europaeus), Bayern, Germany. April
Red snappers / Two-spot red snappers (Lutjanus bohar) mass shoal of fish spawning, Palau, Philippine Sea. A few black and white snappers (Macolor niger) have crawled into the school to eat the eggs
Yellow-flecked glassfrogs (Sachatamia albomaculata) mating pair, Osa Peninsula, Costa Rica
Veragua cross-banded treefrogs (Smilisca sordida) mating pair at night, Osa Peninsula, Costa Rica
Green mandarinfish (Synchiropus splendidus) pair courting and mating at sunset, Sulu Sea, Philippines
American elk (Cervus elaphus canadensis) stag sticking tongue out, Yellowstone National Park, Wyoming, USA
Anemone Hermit Crab (Pagurus prideaux) courtship behaviour. Maseline Harbour, Sark, British Channel Islands, September
Velvet Swimming Crab (Necora / Liocarcinus puber) male guarding female. Gouliot Caves, Sark, British Channel Islands, August
Northern gannets {Morus bassanus} pair in courtship display, Great Saltee, Co. Wexford, Ireland, June
Capercaillie (Tetrao urogallus) male displaying and calling at lek, Kamchatka, Far east Russia, May
Long-tailed manakin (Chiroxiphia linearis) perched on lek (display perch), Costa Rica
Sooty Albatross (Phoebetria fusca) group courting on high inland ridge. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic. Endangered
RF - Red deer (Cervus elaphus) stags fighting, Holkham Park, Norfolk, England, UK, September
Black Grouse (Tetrao tetrix) lek with male displaying and females around in winter, Tver, Russia. April
Black Grouse (Tetrao tetrix) males fighting in winter, Tver, Russia. April
Victorias riflebird (Ptiloris victoriae) male displaying in rainforest, Atherton Tablelands, Queensland, Australia, October
Horned puffins (Fratercula corniculata) pair interacting by touching bills while perched on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Mediterranean Monk seal (Monachus monachus) pair courting on rocks on seabed, Deserta Grande, Desertas Islands, Madeira, Portugal, August 2009, Critically endangered species
Male Spanish ibex (Capra pyrenaica) scenting air behind female with tongue, Sierra de Gredos, Spain, November 2008. WWE OUTDOOR EXHIBITION. NOT AVAILABLE FOR GREETING CARDS OR CALENDARS
Pair of Common hermit crabs (Pagurus bernhardus) male carrying female until she leaves her shell and they can mate, Trondheimsfjorden, Norway, February 2009
Grey seal (Halichoerus grypus) mating behaviour, Donna Nook, Lincolnshire, UK, November 2008
Grey seal (Halichoerus grypus) pair, Donna Nook, Lincolnshire, UK, November 2008
European green toad (Bufo viridis) calling, vocal sac inflated, with eyes closed, in a pond (at 2, 711m) Adylsu valley, side valley to Baksan valley and Elbrus, Caucasus, Russia, June 2008
Red-backed shrike (Lanius collurio) pair, male feeding insect to female in courtship, Bulgaria, May 2008
Common roller (Coracias garrulus) female feeding on its courtship gift, Bulgaria, May 2008
Spiny spider crab (Maja brachydactyla / squinado) pair on rock covered with red algae, Lundy Island Marine Conservation Zone, Devon, England, UK, May
European Hares (Lepus europaeus) boxing, female on right. Wales, UK, June. Sequence 1 of 2
Red deer (Cervus elaphus) stag bellowing, rutting season, Roehampton Flats in background, Richmond Park, London, UK, September
Red deer (Cervus elaphus) stag portrait, bellowing with bracken in antlers, rutting season, Bushy Park, London, UK, October
Two Fallow deer (Dama dama) bucks during fight, rutting season, Richmond Park, London, UK, October
Pair of Common cuttlefish (Sepia officinalis), female in foreground and male behind, note circular mating scars on female caused by the males suckers, Babbacombe Bay, Torbay, Devon, UK, May
Common blue damselfly {Enallagma cyathigerum}, mating pair, Little Bradley Ponds, Devon, UK. July. 2020VISION Book Plate
Black grouse (Tetrao tetrix) males fighting at lek site, Cairngorms National Park, Grampian, Scotland, UK, April
Black grouse (Tetrao tetrix) male in flight at lek, Cairngorms NP, Grampian, Scotland, UK, April
Red deer (Cervus elaphus) stag bellowing, Bradgate Park, Leicestershire, England, UK, October
Black grouse (Tetrao tetrix) male displaying at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two Black grouse (tetrao tetrix) males fighting at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two Black grouse (Tetrao tetrix) males displaying at a lek site at dawn, Cairngorms National Park, Scotland, UK, April
Black necked grebe (Podiceps nigricollis) pair at night, performing their courtship dance during the mating season, with street lights reflected in the water. The Netherlands.April 2014
Red-crowned Cranes (Grus japonensis) displaying and calling in snow, Hokkaido, Japan, February
Pair of Rough snout ghostpipefish (Solenostomus paegnius), with female with egg pouch in front of male. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
