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Black grouse (Lyrurus tetrix), male at lek, Finland, April
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Western capercaillie (Tetrao urogallus) Tver, Russia. May
Black Grouse (Tetrao tetrix) displaying at a lek in snow, Tver, Russia. April
Northern gannet (Morus bassanus) pair, courtship display, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, May
Wood pigeon (Columba palumbus) pair preening one another, The Netherlands
Pair of mating soldier beetles (Rhagonycha fulva) on Common ragwort (Senecio jacobaea), Arne RSPB reserve, Dorset, England, UK, August
Red deer (Cervus elaphus) stags fighting during rut, Richmond Park, London, England, UK, October
Red deer (Cervus elaphus) stag chasing another during a fight, rutting season, Bushy Park, London, UK, October
Two Red deer (Cervus elaphus) stags fighting, rutting season, Bushy Park, London, UK, October
Capercailie (Tetrao urogallus) male displaying from branch of Scots pine tree. Cairngorms National Park, Highlands, Scotland, UK, March
Common goldeneye (Bucephala clangula) male performing its mating display. Southern Norway. March
Spotted owlet (Athene brama) pair on a branch, Bharatpur / Keoladeo Ghana National Park, India
Red deer (Cervus elaphus) stag bellowing during rut, at night with full moon, Derbyshire, UK
Red deer (Cervus elaphus) stag bellowing in mist at sunrise, rutting season, Bushy Park, London, UK, October
Red deer (Cervus elaphus) stag thrashing bracken, rutting season, Bushy Park, London, UK, October
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Diver swimming with an aggregation of Cod (Gadus morhua) over a kelp forest. These cod were gathered in early spring off the north coast of Iceland to spawn. Thorshofn, Iceland. North Atlantic Ocean
Red deer (Cervus elaphus) stag roaring at dusk during rutting season, Cheshire, UK October
Guianan Cock-of-the-rock (Rupicola rupicola), during courtship display, Guianan Shield, Suriname
Argus pheasant / Great argus (Argusianus argus) male displaying, Danum Valley forest reserve, Sabah, Borneo, Malaysia
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Mandarinfish (Synchiropus splendidus) pair spawning. The larger male (behind) is lifting the female into the spawning rise on his pectoral fin, the smaller female (foreground) is releasing eggs
Male Carolas Parotia (Parotia carolae) bird of paradise displaying on his display court to five females perched above, Crater Mountain Wildlife Management Area, Eastern Highlands Province
Waved Albatross (Phoebastria irrorata) courtship, Punta Cevallos, Espaola Island, Galapagos Islands, Ecuador, South America. Endemic, Critically endangered
Green-winged macaws (Ara chloroptera) preening each other. Brazil. South America
Red deer (Cervus elaphus) stag thrashing bracken, rutting season, Bushy Park, London, UK, October. 2020VISION Book Plate
Bower of Vogelkop Bowerbird (Amblyornis inornata) with bright decorations. Nov 2004
Annas hummingbird (Calypte anna) male dancing in flight and displaying his bright pink throat feathers to a female, Montana, USA. June
Firefly (Lamprohiza splendidula) light trails of males in forest at dusk, Bavaria, Germany. July
Greater Prairie-Chicken (Tympanuchus cupido) male displaying on lek. Fort Pierre National Grassland, South Dakota, USA. When displaying
Tench (Tinca tinca) during spawning period amongst European watermilfoil (Myriophyllum spicatum), Sion, Valais canton, southwestern Switzerland
Galapagos penguin (Spheniscus mendiculus), pair courting, Isabela Island, Galapagos Islands
Red fox (Vulpes vulpes) pair nuzzling. London, UK. February
Male Black grouse (Tetrao / Lyrurus tetrix) in flight. Kuusamo, Finland, April
Boat-tailed grackle pair (Quiscalus major) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Lawess parotia (Parotia lawesii) male displaying. Papua New Guinea
Pair of Rock beauty angelfish (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of the female during the spawning rise
Kottigehar dancing frog (Micrixalus kottigeharensis), male waving foot and calling, territorial behaviour. Agumbe, Western Ghats, India. Endemic
Red Deer (Cervus elaphus) stags fighting during the rutting season. Bushy Park, London, UK. October
Giant barrel sponge (Xestospongia testudinaria) releases a large cloud of gametes as it spawns on a coral reef, while a Threadfin hawkfish (Cirrhitichthys aprinus) looks
Sockeye salmon (Oncorhynchus nerka) males fighting in river Adams River, British Columbia, Canada. October
Espanola saddelback tortoise (Chelonoidis hoodensis) pair mating, Espanola Island, Galapagos
Pair of courting Egg-eating sea snakes / Turtleheaded sea snakes (Emydocephalus annulatus) New Caledonia, Pacific Ocean
Colorado river toad (Incilius alvarius) pair mating. Sonoran desert, Arizona, USA. August