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Pair of Great crested grebes (Podiceps cristatus) performing the weed dance during courtship, The Netherlands, Europe. March
Pair of Mandarinfish (Synchiropus splendidus) mating, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Female Yellow-footed rock wallaby (Petrogale xanthopus) fending off advances from male (right), Idalia National Park, Queensland, Australia
Red-throated diver (Gavia stellata) pair displaying on the water, Vaala, Finland. May
Pair of Waved albatross (Phoebastria irrorata) courtship, Espanola Island, Galapagos, South America
Silvery grebe (Podiceps occipitalis) attempting to join Hooded grebes (Podiceps gallardoi) during a breeding display, Patagonia National Park, Santa Cruz, Argentina. Cropped
Pair of Leopard tortoises (Stigmochelys pardalis) mating, Karoo, South Africa
White stork (Ciconia ciconia) pair displaying on their nest built on the chimneys of Knepp Castle, Sussex, UK, June
Northern gannets (Morus bassanus) pair displaying amongst campion, Bempton Cliffs, Yorkshire, UK, May
Pair of mating Big blue octopuses (Octopus cyanea) where the male, on the right, is fertilising the female using its fertilising tentacle. Indian ocean, Maldives
Gentoo penguins (Pygoscelis papua) pair, Sea Lion Island, Falkland Islands
Black-browed albatross (Thalassarche melanophris) pair in courtship. Falkland Islands
Waved albatross (Phoebastria irrorata), pair mutual preening in courtship, Espanola Island, Galapagos Islands
Kingfisher (Alcedo atthis) male passing fish to female, courtship behaviour, Lorraine, France, March
Pink salmon (Oncorhynchus gorbuscha) migrate up river, Vancouver Island, Canada
Bilingual frog (Crinia bilingua) adult male calling, Adelaide River, Northern Territory, Australia, January
Stag beetle (Lucanus cervus) males fighting at dusk in oak wood, the Netherlands, June
Two Black-browed albatross (Thalassarche melanophris) rub their bills together - part of a bonding ritual. West Point Island, The Falkland Islands
Aerial view of Lesser flamingos (Phoeniconaias minor), mating display Lake Magadi, Rift Valley, Kenya
Arctic tern (Sterna paradisaea) pair mating, on rocks on beach. Hailuoto, Finland. May
Red deer (Cervus elaphus) stags fighting during the rutting season, Bushy Park, London, UK October
White-lipped deer (Cervus albirostris) walking on the Tibetan plateau, Serxu, Garze Prefecture, Sichuan Province, China
Clearfin lionfish (Pterois radiata) mating. The female (left) has just released a raft of eggs, which is visible between her tail and the red face of the male
Andean Cock-of-the-rock male (Rupicola peruvianus) at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
African grey hornbill (Lophoceros nasutus) calling in display, perched on tree snag. Janjanbureh, Gambia
Azure / common coenagrion damselflies (Coenagrion puella) pair mating, Meeth Quarry, Devon, UK. July
Sockeye salmon (Oncorhynchus nerka) migration, Adams River, British Columbia, Canada. October
Galapagos hawk (Buteo galapagoensis) landing on mating pair of Galapagos giant tortoise (Chelonoidis vandenburgh) Alcedo Volcano, Isabela Island, Galapagos
Alcedo giant tortoise (Chelonoidis vandenburghi) pair mating, Alcedo Volcano, Isabela Island, Galapagos
Waved albatrosses (Phoebastria irrorata) courting, Punta Suarez, Espanola Island, Galapagos
Alcedo giant tortoise (Chelonoidis vandenburghi) male mating with female while squabbling with another tortoise. Alcedo Volcano, Isabela Island, Galapagos
California least terns (Sternula antillarum browni) courtship behaviour, Bolsa Chica Ecological Reserve, California, USA June
Common Merganser (Mergus merganser) female with ducklings, in threat posture at the approach of a second female, Lansing, New York, USA
Leopard (Panthera pardus) pair mating behaviour, Okavango Delta, Botswana
Black Grouse (Lyrurus tetrix) fighting on icem Finland. March
Eurasian rollers (Coracias garrulus) courting on branch, Hungary. June
Small emerald spreadwing (Lestes virens) pair mating. Klein Schietveld, Brasschaat, Belgium. August
Blue-tailed damselfly (Ischnura elegans) pair mating on stem. Brasschaat, Belgium. August
King cormorants (Phalacrocorax atriceps) in courtship behaviour at their nest site. Saunders Island, Falkland Islands
Red deer (Cervus elaphus) stag bellowing during rut in grassland, doe in background
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
Amur / Siberian tiger (Panthera tigris altaica) pair in courtship, in snow. Captive in tiger park, Heilongjiang Province, China. January
Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef
Fallow deer (Dama dama) herd at dawn, Holkham, Norfolk, England, UK, October
Red deer (Cervus elaphus) stag roaring during the rut, Leicestershire, UK, October. Non-ex
Red deer {Cervus elaphus} adult male in autumnal bracken during the October rut. Leicestershire, UK
Waved albatross (Phoebastria irrorata) pair in courtship display at nest site, Punta Suarez, Espanola Island, Galapagos
Black-browed albatross pair (Thalassarche melanophris) mutual grooming during copulation. New Island, Falkland Islands, November
