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People watching the Fissure eruption of Chico volcano into 9km diameter caldera. Sierra Negra volcano, Isabela Island, Galapagos, Ecuador. October 2005
Lesser horseshoe bats (Rhinolophus hipposideros) in magnesium mine, Shropshire, England, UK, April
King penguins (Aptenodytes patagonicus) walking in line on a windy beach. Sanders Island, Falklands. November
English oak (Quercus robur) ripening acorns, Dorset, UK September
Sulphur tuft mushrooms (Hypholoma fasciculare), Hampstead Heath, England, UK. September
Turnstones (Arenaria interpres) and Purple sandpipers (Calidris maritima) over wintering on the beaches of the Orkney Isles, Scotland, UK, April
Orca (Orcinus orca) surfacing, Shetland, Scotland, UK, August
Gannet (Morus bassanus) diving for fish, Noss, Shetland Islands, Scotland, UK, August
Group of Tissue moths (Triphosa dubitata) hibernating in a limestone cave. Peak District National Park, Derbyshire, UK. November
Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico
White-fronted goose (Anser albifrons) and Taiga bean goose (Anser fabalis rossicus) flocks in flight and on water. Latvia. April
Wirehaired daschund, Staffordshire bull terrier and Hungarian viszla in autumn colours, Wiltshire, UK
Black skimmers (Rynchops niger), three in non-breeding plumage on beach, Fort De Soto Park, St. Petersburg, Florida, USA. January. Digitally altered to remove distractions
White pelicans (Pelecanus erythrorhynchos) at Bear River Migratory Bird Refuge with an approaching storm in May, Utah, USA, May
Red-breasted Mergansers (Mergus serrator) male in foreground performing courtship display to female in background (left), Aurora, New York, USA, April
Wood Ducks (Aix sponsa), females behave aggressively toward a male (second from right) that is trying to join them on a floating log, autumn, New York, USA, October
Wilsons Phalaropes (Phalaropus tricolor) flock flying over tufa formations at Mono Lake in late summer, with reflection, California, USA. July 2013
Avian feeding frenzy with a flock of White Pelicans (Pelecanus erythrorhynchos) swimming along a channel chasing school of fish
Mixed Blackbird flock, mostly Red-winged Blackbirds (Agelaius phoeniceus), in flight against backdrop of autumn foliage in November, Bosque Del Apache National Wildlife Refuge, New Mexico, USA
Snow buntings (Plectrophenax nivalis) flock in flight during a snowstorm, New York, USA, January
Snow Geese (Chen caerulescens) large flock in pool at Bosque Del Apache National Wildlife Refuge with setting full moon at sunrise in December, New Mexico, USA. December
Black-bellied whistling-ducks (Dendrocygna autumnalis), group perched amid Spanish moss on bald cypress tree, Lakeland, Florida, USA, January
European otter (Lutra lutra) group with one shaking off water, Kiskunsagi National Park, Hungary, October
Guanaco (Lama guanicoe) grazing on grassland plain, Patagonia, Chile. Novmeber
Llama (Lama glama) herd, Laguna colorada. Altiplano, Bolivia
Vicuna (Vicugna vicugna), five running across salt flat with mountain in background. Salar de Ascotan, Chile. September 2018
Griffon vulture (Gyps fulvus) landing on coast next to flock. Cantabria, Spain, March
Three French Bulldog puppies, age 6 weeks, sitting in a row
Sergeant major (Abudefduf sp) schooling in mangroves of Isabela Island, Galapagos
Red deer (Cervus elaphus) herd during rut in morning light. Derbyshire, England, UK. October
Marsh fritillary (Euphydryas aurinia), group feeding on Marsh thistle (Cirsium palustre). Lincolnshire, England, UK
Green-crowned brilliant (Heliodoxa jacula), three feeding on Lobster-claw (Heliconia sp) flower in rain. Costa Rica
Black-headed gull (Chroicocephalus ridibundus) flock flying over a frozen lake in falling snow, Wiltshire, UK, March
Greater flamingos (Phoenicopterus roseus) flock, Sado Estuary, Portugal. January
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Pack of wolves (Canis lupus). Yellowstone National Park, Wyoming, USA. January
Thomsons gazelle (Eudorcas thomsonii) herd. Serengeti National Park, Serengeti, Tanzania
Group of Swan geese (Anser cygnoides) on grassland, Inner Mongolia, China
Piraputanga, (Brycon hilarii) reflected on the water surface, Aquario Natural, Bonito, Mato Grosso do Sul, Brazil
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Bohar snapper (Lutjanus bohar) school at Ras Mohammed, Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea
Masai giraffe (Giraffa camelopardalis tippelskirchi) male, females and calf standing close together, Masai Mara Game Reserve, Kenya
Thomsons gazelle (Eudorcas thomsonii) male and two females standing in rainstorm, Masai Mara Game Reserve, Kenya
Whale Shark (Rhincodon typus) and Golden trevally (Gnathanodon speciosus) Cenderawasih Bay, West Papua, Indonesia
Killer whale (Orcinus orca) hunting inside herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Cockroaches (Periplaneta americana) on wall of Gomantong Cave. Sabah, Borneo. Malaysia
Galapagos racer snakes (Pseudalsophis biserialis) group alert watching for prey on beach, Galapagos, June. Endangered species. Taken on location for BBC tv Blue Planet 2 series
Flock of Mallard ducks (Anas platyrhynchos) taking off, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. December
