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Striped mackerel (Rastrelliger kanagurta) with mouths wide open as they swim through the water, filtering zoopankton with their gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Cichlids (Cichlidae) in Lake Malawi, Malawi, November. Photographed for The Freshwater Project
Red Campion (Silene dioica) against shaded background in deciduous woodland, Berwickshire, Scotland, June
Monarch butterfly (Danaus plexippus), in wintering from November to March in Oyamel pine forests (Abies religiosa) Monarch Butterfly Biosphere Reserve, Mexico
Siberian ibex (Capra sibirica) herd, Altai Mountains, Gobi Desert, Mongolia. November
Striped mackerel (Rastrelliger kanagurta) shoal swimming with open mouths, filtering for zooplankton. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Tawny nurse shark (Nebrius ferrugineus) group, individuals circling each other at dusk. Vavuu Atoll, Maldives. Indian Ocean
Spotted hyena (Crocuta crocuta), cubs together by den, Masai-Mara Game Reserve, Kenya
Svalbard reindeer (Rangifer tarandus platyrhynchus) mother and calves, Svalbard, Norway, February
Acrobat ant (Crematogaster sp) tending aphids on white oak (Quercus sp) Washington State Park, Pennsylvania, USA September
Aggregation of thousands of Spider crabs (Leptomithrax gaimardii) for moulting, South Australia Basin, Australia. Pacific Ocean
Northern gannet (Morus bassanus) in flight over colony, Quebec, Canada, August
Group of California sea lions (Zalophus californianus) in a kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
Canada goose (Branta canadensis) adult swimming leading five goslings, Ithaca, New York, USA
Juvenile male Peregrine falcon (Falco peregrinus) in flight chasing his parent who has Feral pigeon (Columba livia) kill in claws, Bristol, England, UK, June
RF- Small group of Pink-footed geese (Anser brachyrhynchus) in flight at dawn, The Wash Estuary, Norfolk, England, UK, October 2011
Chilean flamingos (Phoenicopterus chilensis) group of three in flight, Lago Amarga, Torres del Paine National Park, Patagonia, Chile
Caribbean Flamingo (Phoenicopterus ruber) chick group walking around the breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June
Aerial view of Caribbean Flamingo (Phoenicopterus ruber) breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May
Three kittens with different black and white markings, sitting together, against white background
Flock of White ibis (Eudocimus albus) in breeding plumage, at rookery on water sFlock of White ibis (Eudocimus albus) in breeding plumage, at rookery on waters edge of the mangrove-covered island, Tampa Bay, Florida, USA, March
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by pilotfish (Naucrates ductor) as it cruises beneath the surface of the Red Sea in the late afternoon, close to Little Brother Island
Female Capybara (Hydrochoerus hydrochaeris) swimming in line with young after escaping a Jaguar attack (Panthera onca palustris) in a lagoon off the Paraguay River
Cavalier King Charles Spaniel, puppies with blenheim colouration, resting in straw, age 8 weeks
Group of California sea lions (Zalophus californianus) swimming in kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
Meerkats (Suricata suricatta) standing alert, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Domestic dog, Dogo Argentino, three puppies playing with stick, France
Ice stalagmites / ice spikes formation, Lake Baikal, Siberia, Russia, March
Snorkeller with shoal of Piraputanga (Brycon hilarii), Rio Olha d Agua at Cabeceira do Prata, Bonito, Mato Grosso do Sul, Brazil
Male Ostrich (Struthio camelus) protecting chicks from the sun with its wings, Samburu National Reserve, Kenya, Africa
Female Grizzly bear (Ursus arctos horribilis) standing up, with two cubs nearby, Khutzeymateen Grizzly Bear Sanctuary, British Columbia, Canada, June
Alpaca Herd on Norfolk Farm, winter
Large group of Cavalier King Charles Spaniel, puppies, three of which have tricolour coat and the rest blenheim
Fresh Wych elm (Ulmus glabra) leaves in spring. Dorset, UK May
Grey-cheeked parakeets (Brotogeris pyrrhoptera) perched and grooming on a branch. Guayaquil, Guayas, Ecuador
Raja Brookes Birdwing Butterfly (Trogonoptera brookiana), Borneo
Black-backed jackal (Canis mesomelas) three cubs at the den, Masai-Mara Game Reserve, Kenya
Canada geese (Branta canadensis) flock in flight, Moray Firth, Highlands, Scotland. June
RF - Four colourful Red-and-green macaws or Green-winged macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
Barn owl (Tyto alba) four chicks in nest, La Pampa, Argentina
Great horned owl (Bubo virginianus) chicks. Captive bred, occurs in the Americas
Tawny owl (Strix aluco) with three chicks, against white background
Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by a group of Pilotfish (Naucrates ductor). Rocky Island, Egypt. Red Sea
School of large Atlantic bluefin tuna (Thunnus thynnus) captive in growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
School of Longnose parrotfish (Hipposcarus harid) swarming over reef. Ras Mohammed Marine Park, Sinai, Egypt. Red Sea