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Galapagos racer snakes (Pseudalsophis biserialis) group alert watching for prey on beach, Galapagos, June. Endangered species. Taken on location for BBC tv Blue Planet 2 series
Flock of Mallard ducks (Anas platyrhynchos) taking off, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. December
Blackcap (Sylvia atricapilla) female and two males huddling together for warmth before going to sleep, Finland, May
Common redpoll (Acanthis flammea), flock feeding, Finland, January
Southern Rockhopper penguin (Eudyptes chrysocome) colony, Kidney Island, Falkland Islands, October. Vulnerable species
Gentoo penguin (Pygoscelis papua) walking back to the colony, Sea Lion Island, Falkland Islands, October
Fallow deer (Dama dama) at night, Gyulaj, Hungary Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Meerkats (Suricata suricatta), Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Eastern Ring-tailed Vontsira (Galidia elegans elegans) (endemic family Eupleridae) foraging on forest floor. Marojejy National Park, north east Madagascar
Freshwater isopods (Acanthogammarus victorii) feeding on seabed, Lake Baikal, Siberia, Russia
Indian wild ass (Equus hemionus khur), group standing together, Little Rann of Kutch, Gujarat, India
Starling (Sturnus vulgaris), eating fruit of Japanese Persimmon (Diospyros kaki), Provence, France
Southern elephant seal (Mirounga leonina), fight between two males, Saint Andrew, South Georgia
Series of Bald eagle (Haliaeetus leucocephalus) in flight, Homer, Alaska, USA, March. Digital composite
Puna / James flamingo (Phoenicoparrus jamesi) flock on Laguna Colorado, Reserva Eduardo Avaroa, Altiplano, Bolivia
European Chubb (Squalius / Leuciscus cephalus) in cave, Plitvice Lakes National Park, Croatia. November
School of Munks devil ray (Mobula munkiana), feeding on plankton at night, photographed on a long exposure, Espiritu Santo Island, Sea of Cortez, Baja California, Mexico, East Pacific Ocean
Grey wolf (Canis lupus) mother and two month old cubs, pups begging for food by licking mothers mouth, captive
Red Deer (Cervus elaphus) herd in forest in snow, Cairngorms National Park, Scotland, UK. December
Polar bear (Ursus maritimus) sow with two cubs walking past a pile of Bowhead whale (Balaena mysticetus) bones, Kaktovik, Barter Island, 1002 Area, Arctic National Wildlife Refuge, North Slope
Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Pink-footed goose (Anser brachyrhynchus) flock flying in mist, Cromarty Firth, Highlands, Scotland, UK April
Curlew (Numenius arquata) roosting on saltmarsh, Cromarty Firth, Scotland, UK April
Chilean devil rays / Mobulas (Mobula tarapacana) offshore Santa Maria, Azores, Portugal, Atlantic Ocean
Greater flamingos (Phoenicopterus roseus) flock reflected, feeding in water, Pont de Gau Ornithological Park, Camargue, France, March
Leaf-cutter ants (Atta cephalotes) carrying pieces of leaf that they have harvested back to their underground fungus garden in their nest, Osa Peninsula, Costa Rica
Plankton at night, Palau, Philippine Sea
Army ants (Eciton sp. ) Costa Rica. February 2015Army ants (Eciton sp.) Costa Rica. February 2015
Alpine ibex (Capra ibex) group posing on a rocky mountain side on misty day, Gran Paradiso National Park, Italy. November
Killer whale (Orcinus orca) adult male surfacing at dusk surrounded by birds, who watch for herrings (Clupea harengus) left after they have fed, Andenes, Andoya island, North Atlantic Ocean, Norway
Banded demoiselle (Calopteryx splendens) males resting on grass, Netherlands, August
Great white egrets (Casmerodius albus) reflected in Pulicat Lake, Tamil Nadu, India, January 2013
RF - Red-and-green macaw (Ara chloropterus) flock feeding and flying. Wall of clay lick, Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
RF- Domestic pig, hybrid large white sow and piglets in sty, UK, September 2010
Big-eye or Goggle-eye (Priacanthus hamrur) and Crown squirrelfish (Sargocentron diadema) on wreck colonised by corals, Red Sea, Egypt. January
Big-eye fish (Priacanthus hamrur). Egypt, Red Sea
Chocolate chip sea star o (Protoreaster nodosus) on sandy bottom. Mabul, Malaysia
Antarctic fur seal (Arctocephalus gazella), mother warning off curious juvenile. Southern elephant seal (Mirounga leonina) in background. Gold Harbour, South Georgia. October
King penguin (Aptenodytes patagonicus) colony. Right Whale Bay, South Georgia. September
Razorbill (Alca torda) group, Great Saltee Island, County Wexford, Ireland. June
Candle-snuff fungus (Xylaria hypoxylon), New Forest National Park, Hampshire, England, UK. November
Sociable weaver (Philetairus socius) nest in quiver tree (Aloidendron dichotomum) Quiver tree forest, Keetmanshoop, Namibia
Lions (Panthera leo) five drinking together, Ndutu, Tanzania
Bornean orangutan (Pongo pygmaeus) mother and babies, Tanjung Puting National Park, Borneo-Kalimatan, Indonesia, endangered species
Lionesses (Panthera leo) trying to bring down African buffalo (Syncerus caffer) Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa. Sequence 5 of 5
RF - Leopard (Panthera pardus) mother with cubs, Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Arctic fox (Alopex lagopus) juveniles play fighting, Wrangel Island, Far East Russia, August
Musk ox (Ovibos moschatus) herd, Wrangel Island, Far Eastern Russia, March
