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Turnstones (Arenaria interpres) and Purple sandpipers (Calidris maritima) over wintering on the beaches of the Orkney Isles, Scotland, UK, April
Southern hawker (Aeshna cyanea) dragonfly in flight, Broxwater, Cornwall, UK. July
RF - Southern hawker (Aeshna cyanea) dragonfly in flight, Broxwater, Cornwall, UK. August
Painted lady butterfly (Cynthia cardui) feeding on Verbena flowers in flight, England, UK. August
Egyptian goose (Alopochen aegyptiacus) in flight over Rutland Water, Rutland, UK, August
Common tern (Sterna hirundo) in flight, Norfolk, UK, August
Gannet (Morus bassanus) adult, backlit by the morning sun, in flight. Shetland Islands, Scotland, UK. August
Common / Bandit pipistrelle bat (Pipistrellus pipistrellus) hunting at twilight, Berwickshire, Scotland, August
Kittiwake (Rissa tridactyla) in flight, juvenile. Bempton, Yorkshire, UK, August
Sword-billed hummingbird (Ensifera ensifera) in flight, North-Ecuador, Ecuador
White-fronted goose (Anser albifrons) and Taiga bean goose (Anser fabalis rossicus) flocks in flight and on water. Latvia. April
American avocets (Recurvirostra americana) two during aggressive (territorial) encounter, Bear River Migratory Bird Refuge, Utah, USA, May
Abstract patterns of reflected sky and clouds on the ocean with a silhouetted Red-faced cormorant (Phalarocorax urile) in flight, St. Paul Island, Pribilof Islands, Alaska, USA, July
Great crested flycatcher (Myiarchus crinitus), female alighting at nest cavity entrance carrying pine needles as nest material, wide angle captured with remote camera to show habitat, New York, USA
Tufted puffin (Fratercula cirrhata), adult in breeding plumage flying in preparing to land, St. Paul, Pribilof Islands, Alaska, USA, July
Wilsons Phalaropes (Phalaropus tricolor) flock flying over tufa formations at Mono Lake in late summer, with reflection, California, USA. July 2013
Horned Puffin (Fratercula corniculata) flying over a pattern of kelp and sea foam, photographed from above, St. Paul, Pribilof Islands, Alaska, USA. July
Mixed Blackbird flock, mostly Red-winged Blackbirds (Agelaius phoeniceus), in flight against backdrop of autumn foliage in November, Bosque Del Apache National Wildlife Refuge, New Mexico, USA
Snow buntings (Plectrophenax nivalis) flock in flight during a snowstorm, New York, USA, January
Snowy Egret (Egretta thula) adult in breeding plumage taking flight, Viera Wetlands, Florida, USA, March
Male Red winged blackbird (Agelaius phoeniceus) taking off, New York, USA, May
American Dipper (Cinclus mexicanus) in flight over white water rapids. Lee Vining Creek, Mono Lake Basin, California, USA, April
Griffon vulture (Gyps fulvus) landing on coast next to flock. Cantabria, Spain, March
Rivolis hummingbird (Eugenes fulgens) nectaring on Fuchsia (Fuchsia sp) flower. Costa Rica
Leaf-nosed bat (Phyllostomidae sp) flying towards Banana (Musa sp) flower to feed. Costa Rica
Black-headed gull (Chroicocephalus ridibundus) flock flying over a frozen lake in falling snow, Wiltshire, UK, March
Black-legged kittiwake (Rissa tridactyla) in flight over Newcastle city centre in a blizzard. Newcastle, UK. March
Garden bumblebee (Bombus hortorum) flying to Hedge woundwort (Stachys sylvatica) Monmouthshire, Wales, UK, July
Common Swift (Apus apus) in flight reflected in water, Norfolk, England, UK. July
Little bittern (Ixobrychus minutus) watching Great warbler (Acrocephalus arundinaceus) with prey, Lake Csaj, Kiskunsagi National Park, Hungary
Two coloured mason bee (Osmia bicolor) bee that nests in old snail shells and then carries bits of twig or vegetation to cover the shell, Buckinghamshire, England, UK, June
Copper-rumped hummingbird (Saucerottia tobaci) feeding, Tobago, West Indies
Flock of Mallard ducks (Anas platyrhynchos) taking off, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. December
Common raven (Corvus corax) two adults in flight, Hornoya, Norway, April
Eurasian griffon vulture (Gyps fulvus) in flight, Cevennes, France, March 2016
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
Brown booby (Sula leucogaster), adult diving for fish, Los Roques, Venezuela
Series of Bald eagle (Haliaeetus leucocephalus) in flight, Homer, Alaska, USA, March. Digital composite
Rock ptarmigan (Lagopus muta) in flight, Utsjoki, Finland, April
Blue cheeked bee eater (Merops persicus) two squabbling, Oman, April
Siskin (Carduelis spinus) male in flight preparing to land, Scotland, UK, May
Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Pink-footed goose (Anser brachyrhynchus) flock flying in mist, Cromarty Firth, Highlands, Scotland, UK April
Hoopoe (Upupa epops) male in flight, Hungary May
Green lacewing (Abachrysa eureka) in flight, , Tuscaloosa County, Alabama, USA Controlled conditions. May
Harlequin ladybird (Harmonia axyridis) flying, Tuscaloosa County, Alabama, USA Controlled conditions. January
Red wasp (Polistes carolina) in flight Lamar County, Texas, USA Controlled conditions. August
Hawk owl (Surnia ulula) takingn off from powder snow, possibly with prey in talons, Kuusamo Finland February
