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Japanese dwarf flying squirrel (Pteromys volans orii) gliding through forest canopy during light snowfall. Hokkaido, Japan. February
Wood pigeon (Columba palumbus) flying with nesting material in beak, Lorraine, France, April
Great grey owl (Strix nebulosa) hunting over snow, woodland in background. Kuusamo, Finland. February
Great grey owl (Strix nebulosa) close up hunting in snow. Kuhmo, Finland. February
Osprey (Pandion haliaetus) carrying fish above river in Taipei, Taiwan
African fish eagle (Haliaeetus vocifer) attacking a female Saddle-billed Stork (Ephippiorhyncus senegalensis )trying to steal a fish it has just caught. Liuwa Plains National Park, Zambia
Golden eagle (Aquila chrysaetos) Sodankyla, Lokka, Finland, April
Shoebill stork flying (Balaeniceps rex) in the swamps of Mabamba, Lake Victoria, Uganda
Sparrowhawk (Accipiter nisus) adult female, Norway, October
Grey-headed flying-fox (Pteropus poliocephalus) spotting and flaring wings to come in to land on a branch. Yarra Bend Park, Kew, Victoria, Australia. ÔÇ¿November 2019
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Canada Goose (Branta canadensis) in flight. Gilbert Water Ranch, Arizona, USA. January
Hummingbird hawk-moth (Macroglossum stellatarum) nectaring on Loofah (Luffa sp) flower. Xinjiang, China
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Andean condor (Vultur gryphus), Male, Antisanilla Reserve, Ecuador
Blue-footed booby (Sula nebouxii) landing, Santa Cruz Island, Galapagos, Ecuador
Violet-tailed sylph hummingbird (Aglaiocercus coelestis) in flight, about to land. Ecuador, May
Small flock of Lapwings (Vanellus vanellus) flying in to rest after migration. Cresswell Pond, Druridge Bay, Northumberland, England, UK, September
Common buzzard (Buteo buteo) landing on prey, wings open, Scotland, UK, September 2016
Southern hawker (Aeshna cyanea) dragonfly in flight, Broxwater, Cornwall, UK. August
Garden bumblebee (Bombus hortorum) taking off from Knapweed, England, UK, August
Canada geese (Branta canadensis) in flight at Abbotsbury Swannery, Abbotsbury, Dorset, England, UK. August 2014
Sparkling Violet-ear (Colibri coruscans) hummingbird in flight, Mindo cloud forest ecosystem, Ecuador, July
Golden eagle (Aquila chrysaetos) in flight over snow. Kalvtrask, Vasterbotten, Lapland, Sweden. January
Arctic terns (Sterna paradisaea), two in flight over nesting colony in field of buttercups (Ranunculus sp.), Keflavik, Iceland. Digital canvas expansion
Cinnamon teal (Anas cyanoptera), male taking flight from water, Orange County, California, USA, April
Huge flock of White ibis (Eudocimus albus) in breeding plumage, flying to their mangrove-covered island rookery, Tampa Bay, Florida, USA, March
Violet sabrewing hummingbird (Campylopterus hemileucurus) nectaring on Orchid. Costa Rica
Glowworm beetle (Phengodes sp. ) male flying, Tuscaloosa County, Alabama, USA Controlled conditionsGlowworm beetle (Phengodes sp.) male flying, Tuscaloosa County, Alabama, USA Controlled conditions. July
Small scissor bee (Chelostoma campanularum) In flight approaching garden bee hotel carrying mud to seal nest tube, Hertfordshire, England, UK, June
Blue-footed booby (Sula nebouxii), six diving into sea. Espumilla Beach, Santiago Island, Galapagos
Red bat (Lasiurus borealis) female flying; shots taken with high speed flash San Saba County, Texas, USA. Controlled conditions. July
Blackbird (Turdus merula) in flight to feed on berries, Helsinki, Finland. November
Great grey owl (Strix nebulosa) taking off from snowy ground, Kuusamo, Finland. March
Flock of Tree sparrows (Passer montanus) in flight, with a Greenfinch (Chloris chloris) blurred motion, Pacsmag, Hungary
Long-tailed duck (Clangula hyemalis), males in flight, Finland, April
Kingfisher (Alcedo atthis) male, after diving, taking off from water with fish, a Common Roach (Rutilus rutilus) Lorraine, France, August
Common swift (Apus apus) group flying overhead, Bradford-on-Avon, Wiltshire, UK, May
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Scarlet ibis (Eudocimus ruber), in flight, Coro, Venezuela
Great Frigatebird (Fregata minor) arriving at nest with chick, San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Stellers Sea Eagle (Haliaeetus pelagicus) with prey in beak in mid-air, Hokkaido, Japan, February
Lammergeier / Bearded vulture (Gypaetus barbatus) flying, Leukerbad, Valais, Switzerland, February. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Green june beetle (Cotinis nitida) in flight Williamson County, Texas, USA Controlled conditions. October
Noctule Bat in flight showing teeth, Germany
White tailed sea eagle (Haliaeetus albicilla) perched on tree stump with fish. Hooded crow (Corvus cornix) flying in to attack in background. Danube Delta, Romania. May
Shag (Phalacrocorax aristotelis) in flight over sea. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Razorbill (Alca torda) landing, Great Saltee Island, County Wexford, Ireland, June
