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Pin-tailed whydah (Vidua macroura) male in display flight. Allahein River, Gambia
RF - Atlantic Puffins (Fratercula arctica) flying on to the cliff top and gathering around burrows. Isle of Lunga, Treshnish Isles, Isle of Mull, Scotland
Gila woodpecker (Melanerpes uropygialis), emerging from nest in Saguaro cactus, Arizona, USA. July
Marsh harrier (Circus aeruginosus) female hunting among a panicked flock of Common teal (Anas crecca) in flight over marshland, RSPB Greylake Nature Reserve, Somerset, UK, December
Atlantic puffin (Fratercula arctica) taking off, Machias Seal Island, Maine, USA, June
Common tern (Sterna hirundo) close up in flight, Tenerife, Canary Islands
Galapagos hawk (Buteo galapagoensis) flying, carrying an iguana carcass, Punta Espinosa, Fernandina Island, Galapagos
Short-eared owl (Asio flammeus) in flight, Santa Fe Island, Galapagos
Buff-tailed bumblebee (Bombus terrestris) queen about to land at her nest burrow in a garden lawn with loaded pollen sacs to provision grubs that will become future workers for her colony, Wiltshire
Firefly (Lamprohiza splendidula) light trails of males in forest, Bavaria, Germany. July
Golden eagle (Aquila chrysaetus) landing in snow, Kuusamo, Finland, December
Dipper (Cinclus cinclus) taking off from water, Kuusamo, Finland, February
Bullfinch (Pyrrhula pyrrhula) in flight, Kuusamo, Finland, March
Black Woodpecker male (Dryocopus martius) Finland, January. One branch removed
Great white pelican (Pelecanus onocrotalus) flying over water, Danube Delta, Romania. July
Cuckoo (Cuculus canorus) in flight, Germany, April. May
Great horned owl (Bubo virginianus) in flight. Saskatchewan, Canada, August
Red-and-green macaw (Ara chloropterus) flock feeding at wall of clay lick, flying and perched in trees. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Greater flamingo (Phoenicopterus roseus) legs of bird in flight, Pont Du Gau Park, Camargue, France
Scopolis shearwater (Calonectris diomedea) in flight. Tenerife, Canary Islands
Brunnichs guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018
Barn Owl (Tyto alba) landing on gravestone, captive, digital composite, UK
Magnificent frigatebird (Fregata magnificens) in flight over sea between Isabela Island and Fernandina Island, Galapagos
California condor (Gymnogyps californianus) in flight, wings radio tagged. California condor recovery program, Sierra de San Pedro Martir National Park, Baja California Peninsula, Mexico. 2011
Purple throated mountain gem hummingbird (Lampornis calolaemus) hovering as it nectars on Porterweed (Stachytarpheta frantzii). Costa Rica
Rhinoceros hornbills (Buceros rhinoceros) pair flying over the canopy with mist hanging over lowland Dipterocarp rainforest in heart of Danum Valley, Sabah, Borneo (digitally modified image)
Short-toed Eagle (circaetus gallicus) flying amongst Stone Pines (Pinus pinea). Coto Donyana, Andalusia, Spain. 2007
Green winged macaw (Ara chloroptera) pair flying, Chapada dos Guimaraes, Mato Grosso, Brazil
Snowy owl (Bubo scandiacus) Female taking off the winter tree, Quebec, Canada
Waxwings (Bombycilla garrulus) feeding on snow covered Rowan (Sorbus) berries, whilst in flight, Kuusamo, Finland, Scandinavia, January. Magic Moments book plate
Wandering albatross (Diomedea exulans) flying on the open ocean, Drake passage, Antarctic Peninsula, Antarctica
Barn owl (Tyto alba) hovering whilst hunting in flight. Suffolk, UK. June, Cropped
Early bumblebee (Bombus pratorum), in flight, Monmouthshire, Wales, UK. May
Brent geese (Branta bernicla) flying with moon, Titchwell, Norfolk, UK, December
Common crane (Grus grus) flock in flight, Lac du Der, France, November
Flock of Pink footed geese (Anser brachyrhynchus) in flight at dusk, Norfolk, England, UK, October
Common / Eurasian cranes (Grus grus) flying from roost site at sunrise, silhouetted against dawn sky, autumn migration period, Rugen-Bock-Region, Mecklenberg-Vorpommern, Germany, October 2009
Common Crane (Grus grus) juvenile and two adult calling, displaying, in harvested corn fields in the Rugen-Bock-Region in autumn before they continue their annual journey to the south
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
Swallow-tailed gull (Creagrus furcatus) in flight, Plazas Islands, Galapagos
Galapagos hawk (Buteo galapagoensis) landing, Galapagos, Ecuador. Vulnerable species
Galapagos hawk (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Captive reared juvenile White stork (Ciconia ciconia) in flight over the Knepp Estate soon after release, Sussex, UK, August 2019
Flock of Oystercatchers (Haematopus ostralegus) and Knot (Calidris canuta) in flight and on tidal shallows. The Wash Estuary, Norfolk, September
Barn Swallow (Hirundo rustica) feeding a fledgling on a wire. Perthshire, Scotland, September
Oystercatcher (Haematopus ostralegus) flock in flight, Liverpool Bay, Lancashire, UK, September
Group of Greylag geese (Anser anser) flying in the half light before dawn, Snettisham RSPB reserve, Norfolk, UK, September
Turnstones (Arenaria interpres) and Purple sandpipers (Calidris maritima) over wintering on the beaches of the Orkney Isles, Scotland, UK, April