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Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Silfra Canyon - deep fault filled with fresh water in the rift valley between the Eurasian and American tectonic plates, at Thingvellir National Park, Iceland. June 2014
A split level digital composite showing a Basking shark (Ceterhinus maximus) feeding on plankton around St Michaels Mount, Cornwall, UK. June
Polar bear (Ursus maritimus) walking on Champ Island glacier above the sea, Franz Josef Land, Russian Arctic. Highly commended in Animals in their Environment Category of the Wildlife Photographer of
A jogger on the towpath of the Canal du Midi near Castelnaudary, Languedoc-Rousillon, France. Septeber 2011
Underwater river scene with freshwater plants and Tetra fish, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul
Duckling swimming on water surface, split level, captive, UK
Puffin (Fratercula arctica) swimming underwater, Farne Islands, Northumberland, UK, July
Lowland gorilla adult male silverback in zoo, USA
Mountain Gorilla (Gorilla beringei beringei) Sabyinyo Group, Silverback in meadow, Volcanoes National Park, Rwanda, East Africa
Coopers hawk {Accipiter cooperii} on Saguaro cactus skeleton, Arizona, USA
Golden retriever sitting on boat beside water, USA
Black crowned-crane (Balearica pavonina) foraging in wetland area, Masai Mara, Kenya
Western red cedar tree (Thuja plicata) detail of bark, The Big Tree Trail, Meares Island, Clayoquot Sound, Vancouver Island, British Columbia, Canada
Red Fox (Vulpes vulpes) walking along a fallen trunk in misty forest. Black Forest, Germany, November
American white & Brown pelican P. occidentalis}, Florida, USA
Portrait of a Black-capped chickadee, (Poecile atricapillus) with white background, Block island, Rhode Island, USA. Bird caught during scientific research
Electric fileclam / Disco clam (Ctenoides ales). West Papua, Indonesia. Indo-West Pacific
Male silverback Eastern lowland gorilla (Gorilla beringei graueri) in the equatorial forest of Kahuzi Biega Park, Democratic Republic of Congo, Africa, March 2009
Silverback male Eastern lowland gorilla (Gorilla beringei graueri) intimidation display by chest beating, equatorial forest of Kahuzi Biega Park, Democratic Republic of Congo, Africa, March 2009
Common toad (Bufo bufo) tadpole swimming seen from below, Belgium, June
Scots pine (Pinus sylvestris) trees reflected in Loch Maree at sunrise with Slioch in background, Wester Ross, Scotland, UK, November 2014
Detail of frost on Western Red Cedar (Thuja plicata). Vancouver Island, Canada, March
Raven (Corvus corax) calling in the snow, Kemijarvi, Finland, February
Black-crowned Central American squirrel monkey (Saimiri oerstedii) pair playing with each other, one with hand over eyes of other, Manuel Antonio National Park, Quepos, Costa Rica
White chinned petrel (Procellaria aequinoctialis) in flight low over the water off Kaikoura, Canterbury, New Zealand
A rare British broadsheet illustration with contemporary hand colouring, drawn and engraved by John Emslie and published by James Reynolds in 1849
Eastern lowland gorilla (Gorilla beringei graueri) silverback named Chimanuka climbing a tree, Kahuzi-Biega National Park, South Kivu Province, Democratic Republic of Congo (DRC)
Western lowland gorilla (Gorilla gorilla gorilla) captive in zoo, native to west Africa
Madagascar green-emerald giant pill millipedes (Zoosphaerium neptunus) Andasibe-Mantadia National Park. Madagascar
Butterfly / Rhinoceros viper (Bitis nasicornis). Captive. Occurs in West and Central Africa
Eddystone Lighthouse marking the dangerous Eddystone Rocks off Rame Head in Cornwall
Young European eel (Anguilla anguilla) elvers, or glass eels, caught during their annual migration up rivers from the Bristol channel, swimming in a large holding tank at UK Glass Eels
Sable (Martes zibellina) in tree, Irkutsk, Siberia, Russia. November
Yellow spotted monitor standing threat display {Varanus panoptes panoptes} Queensland, Australia
Bengal Tiger (Panthera tigris) six month old cub jumping on its mother, Bandhavgarh National Park, India
Steam train on the Heritage Poppy Line from Sheringham to Holt, with Weybourne Mill in background, Norfolk, UK, April
Red fox (Vulpes vulpes) cubs playing, Vosges, France, May
Underwater view of Waterlilies (Nymphaea alba) in a lake. Alps, Ain, France, June
Young urban Red fox (Vulpes vulpes) standing on a wall at night. Bristol, UK, September. Nominated in the Melvita Nature Images Awards competition 2014
Water lilies (Nymphaea alba) lit by the suns rays, seen under water. Alps, Ain, France. June. Highly commended in the the Underwater World category
Illustration of human evolution from left to right Australopithecus afarensis, Australopithecus africanus, Homo erectus and Homo sapiens. Pencil and watercolor painting
Golden retrievers in boat {Canis familiaris} USA
Puffin (Fratercula arctica) on a cliff edge with flowering sea thrift (Armeria maritima) Great Saltee, Saltee Islands, County Wexford, Ireland, June
Young girl in traditional dress. Mursi tribe, Mago National Park. Omo Valley, Ethiopia
Wild horse / mustang in McCullough Peaks, Wyoming, USA - buckskin mare and cremello colt Cremosso
Wild Mustang horses interacting, Pryor mountains, Montana, USA. June
Snow leopard mother (Uncia uncia) with cub, captive, occurs in mountains of central and southern Asia
Two Arctic hares (Lepus arcticus) fighting, Northeast Greenland National Park, Greenland. Veolia Environnement Wildlife Photographer of the Year 2009: Mammal Behaviour category: Runner-up
Bottlenose dolphin (Tursiops truncatus) playing in the waves of an oil tanker, Port Aransas, Texas, USA
Red fox in a snow storm (Vulpes vulpes) Winter winds on Kamchatka can reach 40 meters a second. Kronotskty Zapovednik, Kamchatka, Russia
Barn and wheat field in the Palouse farming area of southeastern Washington, USA, August
Carp (Cyprinus carpio) Aare River, Switzerland 2006
Norwejian Fjord stallion running in the snow, Berthoud, Colorado, USA
Brown trout (Salmo trutta) leaping for a damselfly. (digitally enhanced)
Eurasian crag martin (Ptyonoprogne rupestris) Kawakarpo Mountain, Meri Snow Mountain National Park, Yunnan Province, China. May
Cudbear Lichen (Ochrolechia tartarea) growing on quartzite rock. Cairngorms National Park, Grampian Mountains, Scotland, UK, February
Bamboo lemur (Hapalmur griseus) in tree. Andasibe-Mantadia National Park, Madagascar
Japanese crane (Grus japonensis) displaying, wings in heart shape, Hokkiado, Japan, February
Brown-mantled tamarin / Saddleback tamarin (Saguinus fuscicollis) climbing down tree trunk, Tambopata, Madre de Dios, Peru
Large group of Sunflower sea stars (Asterias / Pycnopodia helianthoides) covering rock, pacific coast, Canada, August
Kaleidoscopic image of Clarks anemonefish (Amphiprion clarkii), Ambon, Indonesia
The Newport Harbor Light (Goat Island Light) standing framed by the giant Newport Bridge, Rhode Island, USA
Pack of European grey wolves standing in snow (Canis lupus) captive. Bayerischerwald National Park, Germany
Fireworks at Peterhead harbour, Scotland, UK, July 2016
Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December
Gayal / Mithun (Bos frontalis) on river bank. Arunachal Pradesh, North East India. November 2014
Kakamega Forest at dawn, Kenya
Snares island crested penguin (Eudyptes robustus) colony on the coast, high angle view, Snares Island, New Zealand
Himalayan honey bees (Apis dorsata laboriosa) on comb, Mount Namjagbarwa, Yarlung Zangbo Grand Canyon National Park, Tibet, China. October
Bat fish juvenile {Halicutaea sp} deep sea
Glowworm beetle (Phengodes sp. ) male flying, Tuscaloosa County, Alabama, USA Controlled conditionsGlowworm beetle (Phengodes sp.) male flying, Tuscaloosa County, Alabama, USA Controlled conditions. July
Common redstart (Phoenicurus phoenicurus) nest with six eggs, Alsace, France
Caribbean reef squid (Sepioteuthis sepioidea) swimming in open water at night, Guadeloupe Island, Mexico. Caribbean
Scots pine (Pinus sylvestris) forest with Bracken (Pteridium aquilinum) ground flora in autumn, Glenfeshie, Cairngorms National Park, Scotland, UK, October
Noctule Bat in flight showing teeth, Germany
Brown bear (Ursus arctos) paw seem from under water, Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August
Chimpanzee (Pan troglodytes) portrait of orphaned juvenile Sophie, sucking piece of grass, Sweetwater Sanctuary, Kenya
Chimpanzee orphan Sophie eating fruit (Pan troglodytes) Sweetwaters Chimpanzee Sanctuary, Kenya
Arctic Tern (Sterna paradisaea) hovering in flight, June, Iceland. Magic Moments book plate
Urban Red fox (Vulpes vulpes) walking past wall with red fox mural / graffiti. North London, England, UK, April
Snowy Owl (Nyctea scandiaca) adult female flying over snow, winter, Europe Captive /Snowy Owl (Nyctea scandiaca) adult female flying over snow, winter, Europe Captive/ Falconers bird, Austria
Young girl wearing traditional headdress, Mursi tribe, Mago National Park. Omo Valley, Ethiopia
The forests of the Modi Khola river valley shrouded in fog. Annapurna Sanctuary, central Nepal, November 2011