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Portrait of Bigfin squid (Sepioteuthis lessoniana) hovering in open water above a coral reef at night. Dampier Strait, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Great crested grebe (Podiceps cristatus) pair performing the weed dance, a spectacular part of their elaborate courtship ritual, Derbyshire, UK, March
Juvenile Andean mouse opossum (Marmosops impavidus) eating a moth, El Oro, Ecuador
Sinbad the male Kakapo (Strigops habroptilus) curiously peering from the bushes during the day. Codfish Island, Stewart Island, New Zealand, January. Critically endangered
Mountain tree shrew (Tupaia montana) feeding on nectar secreted by the endemic Pitcher Plant (Nepenthes kinabaluensis) Montane forests (at 2200m-3000m), slopes of Mt Kinabalu
Leopard (Panthera pardus) hunting Springbok (Antidorcas marsupialis) Etosha, Namibia, Finalist in the Mammals Category of the Wildlife Photographer of the Year 2015
Marine iguana (Amblyrhynchus cristatus) underwater. Fernandina Island. Galapagos, Endemic Species
Platypus {Ornithorhynchus anatinus} swimming underwater Tasmania, Australia. Commposite
Greater horseshoe bat {Rhinolophus ferrumequinum} in flight hunting a moth at night, Germany
RF - Coquerels sifaka (Propithecus coquereli) jumping, Anjajavy Private Reserve, north west Madagascar
Emperor angelfish (Pomacanthus imperator) swimming past soft corals (Dendronephthya sp]), Red Sea, Egypt. January
Great hammerhead shark (Sphyrna mokarran) in shallow water. South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Red-eyed tree frog (Agalychnis callidryas) swallowing a spider. El Arenal, Costa Rica
Magnificent bird of paradise (Diphyllodes magnificus) male, Lower montane highlands near Mount Hagen, Enga Province, Papua New Guinea. Endemic
Bee orchid (Ophrys apifera) in flower. Dorset, UK, June
Three-striped poison frog (Ameerega trivittata) male carrying tadpoles, Panguana Reserve, Huanuco province, Amazon basin, Peru
Black-striped capuchin (Sapajus libidinosus) using rock as a tool to break open palm nut, Parnaiba Headwaters National Park, Piaui, Brazil. August
Brown-throated Three-toed Sloth (Bradypus variegatus) wild sloth in tree at Aviarios Sloth Sanctuary, Costa Rica
Velvet Worm (Peripatus novaezealandiae) known as living fossils, having remained the same for approximately 570 million years, New Zealand, Captive
Purple sea pen (Virgularia gustaviana) on sandy sea bed. Rinca, Indonesia
Argus pheasant / Great argus (Argusianus argus) male displaying, Danum Valley forest reserve, Sabah, Borneo, Malaysia
Malayan colugo {Cynocephalus variegatus} hanging upside-down in tree feeding on algae at night, Danum Valley, Sabah, Borneo, September
Lava Cactus (Brachycereus nesioticus), growing in lava field, coast of Fernandina, Galapagos Islands, Ecuador
Peringueys adder (Bitis peringueyi) sidewinding in Namib desert, Namibia
Common hawker (Aeshna juncea) Brackish Moss NNR, County Armagh, Northern Ireland
Hourglass treefrog (Dendropsophus ebraccatus) male with inflated vocal sac calling at night, Central Caribbean foothills, Costa Rica
Marine iguanas (Amblyrhynchus cristatus) basking on volcanic rock, Punta Espinosa, Fernandina Island, Galapagos Islands, East Pacific Ocean
Wolf Spiders (Pardosa sp. ), male RIGHT waving palps in courtship displayWolf Spiders (Pardosa sp.), male RIGHT waving palps in courtship display. Peak District National Park, Derbyhsire, UK. May
Black skimmer (Rynchops flavirostris) fishing by skimming along water surface, Pantanal, Brazil
Endive Pellia liverwort (Pellia endiviifolia) in centre growing through Common Liverwort (Marchantia polymorpha), the latter bearing cups containing gemmae (used in asexual reproduction)
Nile crocodile (Crocodylus niloticus) making its way down sand bank into to the Rufiji Rivre, Selous Game Reserve, Tanzania
Adult Giant Anteater (Myrmecophaga tridactyla) walking across savannah. Los Llanos, Colombia, South America
Thomsons gazelle (Eudorcas thomsonii) herd with Wildebeest herd visible beyond, Masai-Mara game reserve, Kenya
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at sunrise. Kasanka National Park, Zambia
Short-beaked echidna (Tachyglossus aculeatus) digging, Tasmania, Australia
Golden-headed lion tamarin (Leontopithecus chrysomelas) captive portrait, Apenheul Park, Netherlands
Lyretail anthias / Goldies (Pseudanthias squamipinnis) over coral reef with soft corals (Dendronephthya sp) Egypt, Red Sea
Meadow in flower, with Cork oaks (Quercus suber) in the background, Beja, Portugal, April
Flatworm (Maiazoon orsaki) Raja Ampat, Irian Jaya, West Papua, Indonesia, Pacific Ocean
Sword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devil sSword-billed Hummingbird (Ensifera ensifera) feeding at an Angels or Devils Trumpet Flower (Datura sp.). Yanacocha montane cloud forest, Ecuador
A portrait of a nudibranch (Eubranchus tricolor) on the seabed of a Scottish loch. Loch Carron, Ross and Comarty, Scotland, Great Britain. North East Atlantic Ocean
Two-tailed Pasha butterfly (Charaxes jasius) expanding wings after emerging, Podere Montecucco. Orvieot, Umbria, Italy August. Sequence 9 of 9
Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October
Male Japanese giant salamander (Andria japonicus) emerging from a nest hole in order to breathe, Japan, August
Large Tree Shrew (Tupaia tana) Captive, occurs Borneo and Sumatra
Saucer Magnolia (Magnolia x soulangeana) tree in full flower against blue sky. Stourhead gardens, Wiltshire, UK, April
Light Bulb Sea Squirt (Clavelina lepadiformis). Channel Islands, UK, June
A Large / Pot Bellied Seahorse (Hippocampus abdominalis) perched in nets. Manly, Sydney, New South Wales, Australia, March
Marine springtail / bristletail (Petrobius maritimus) adult resting on stones, Lough Muree, County Clare, Ireland, August
Common / Eurasian crane (Grus grus) flock in flight with snow topped Pyrenees mountains in the background, during southward migration period, Haute Pyrenees, Gascony, France. December 2009
Little ringed plover {Charadrius dubius} nest with four eggs camouflaged on shingle, Lorraine, France
Greater flamingos (Phoenicopterus roseus) mating, in lagoon, Pont Du Gau, Camargue, France, April 2009
Fossil dinosaur {Struthiomimus altus} Royal Tyrrell Museum, Drumheller, Alberta, Canada
Aboriginal rock art, Anbangbang Gallery, Nourlangie Ranges, Kakadu National Park, Northern Territory, Australia
Numbat (Myrmecobius fasciatus) standing on hind legs, Central Australia
Caecilian (Siphonops annulata) amongst leaf litter, Amazon Rainforest. Ecuador
Midwife Toad (Alytes obstetricans) paternal male carrying eggs, S. Yorks, UKMidwife Toad (Alytes obstetricans) paternal male carrying eggs, S.Yorks, UK
Cape cobra {Naja nivea} displaying amongst vegetation, South Africa
Six banded / yellow armadillo {Euphractus sexcinctus} Minas Gerais, Brazil
Black tailed prairie dogs on burrow mound {Cynomys ludocicianus}, Badlands, USA
Apple moss (Bartrimia pomiformis) with spore capsules, Inverness-shire, Scotland, UK
Termite (Isoptera) colony with six-metre high cooling chimney, Bogoria, Kenya
Yellow streaked tenrec (Hemicentetes semispinosum) feeding on a worm, Maroansetra, Madagascar
Coconut / Robber crab (Birgus latros) climbing coconut tree, Aldabra Seychelles