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Humboldt penguin (Spheniscus humboldti) group gathering at landing site. Tilgo Island, La Serena, Chile. Vulnerable species
Elk / Wapiti (Cervus canadensis) herd with young on meadowland against the Grand Tetons, low clouds and the Snake River. Grand Teton National Park, Wyoming, USA, June
Piraputanga fish (Brycon hilarii) in underwater landscape, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul, Brazil
Common starling (Sturnus vulgaris) flocking prior to roosting, Scotland, January 2009
Piraputanga (Brycon hilarii) shoal, Rio Olhio d agua, tributary of Rio da Prata, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil
Red deer (Cervus elaphus) stag bellowing during rut, Derbyshire, UK
Florida Red-bellied Turtle (Pseudemys nelsoni) Rainbow Springs, Rainbow Springs State Park, Florida, USA, March. Taken for the Freshwater Project
Tail of Southern right whale (Eubalaena australis) at sunset, Golfo Nuevo, Peninsula Valdes, UNESCO Natural World Heritage Site, Chubut, Patagonia, Argentina, Atlantic Ocean, October
Volcan Alcedo giant tortoises (Chelonoidis nigra vandenburghi) among steaming fumaroles on western caldera rim, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
Alpine marmot (Marmota marmota), with Mount Grossglockner (3798m) in background, Hohe Tauern National Park, Austria, July
Grizzly bear (Ursus arctos horribilis) in grassland, Yellowstone National Park, Wyoming, USA, October
Marine iguana (Amblyrhynchus cristatus) on coast, Galapagos
Masai giraffes (Giraffa camelopardalis tippelskirchi) in dry season, Masai-Mara Game Reserve, Kenya. March
African elephant (Loxodonta africana) herd silhouetted in distance walking through a storm, Masai-Mara Game Reserve, Kenya. September
Cheetah (Acinonyx jubatus) resting on a termite hill and storm brewing, Masai-Mara Game Reserve, Kenya. March
Cheetah (Acinonyx jubatus) female with cubs sitting in long grass during the dry season, Masai-Mara Game Reserve, Kenya. March
Wildebeest (Connochaetes taurinus) herd walking in a line in the rain at sunset, Masai-Mara Game Reserve, Kenya. September
Male impala (Aepyceros melampus) at sunrise, Masai-Mara Game Reserve, Kenya. September
Cheetah (Acinonyx jubatus) sitting on a termite mound at sunset, Masai-Mara Game Reserve, Kenya. October
Single Male waterbuck (Kobus ellipsiprymnus) in distance silhouetted against big sky and clouds, Masai-Mara Game Reserve, Kenya. October
Topi (Damaliscus korrigum / Damaliscus lunatus) territorial male standing on mound looking across savanna. Masai-Mara game reserve, Kenya
Cheetah (Acinonyx jubatus) female standing below dark storm clouds, Masai-Mara game reserve, Kenya
American Bison (Bison bison) foraging in the snow, Yellowstone National Park, Wyoming, USA. January 2011
Puma / Cougar (Felis concolor) looking out for prey in habitat, Chile. March 2014
Musk ox (Ovibos moschatus) two in habitat, Wrangel Island, Far Eastern Russia, October
Herd of Tibetan Wild Asses / Kiang (Equus kiang) ChangThang, Tso Kar lake, at altitude of 4600m, Ladakh, India, October 2012
Springbok (Antidorcas marsupialis) grazing in front of sand dunes, Namib Naukluft Park, Namibia, June 2006
Grants zebra (Equus burchelli boehmi) herd with large cloud above, Amboseli National Park, Kenya
Leatherback Turtle Hatchling (Dermochelys coriacea) crossing a beach towards the sea at dawn, Cayenne, French Guiana, July
Orcas (Orcinus orca) pod feeding on herring, wide shot showing surrounding landscape, Iceland, January 2013
Kamchatka Brown Bear (Ursus arctos beringianus) on the Pacific coast of Kamchatka. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Red Fox (Vulpes vulpes) in wide landscape with Kronotsky Volcano on the horizon. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Kamchatka Brown Bear (Ursus arctos beringianus) recently risen from hibernation in the caldera of the Uzon Volcano. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, April
African elephants (Loxodonta africana) two bulls sparring at the waterhole in front of the lodge, Ol Donyo Wuas, Mbirikani Group Ranch, Amboseli-Tsavo ecosystem, Chyulu Hills, Kenya
African elephants (Loxodonta africana) large family group on migration, Amboseli National Park, Kenya
Goliath heron (Ardea goliath) near Mopani Dam at dusk, Kruger National Park, Transvaal, South Africa, September
Killer Whale / Orca (Orcinus orca) large bull swimming in Resurrection Bay, Kenai Fjords National Park, outside Seward, Alaska, May
Hoary marmot (Marmota caligata) adult overlooking Exit Glacier in Kenai Fjords National Park, South Central Alaska, USA
Grey heron (Ardea cinerea) beneath waterfall. Ambleside, Lake District, UK, November. Highly commended, Habitat category, British Wildlife Photography Awards (BWPA) competition 2012
Masai giraffe (Giraffa camelopardalis) and her youngster on the savanna landscape at dawn, Serengeti National Park, Tanzania
Blacktip reef sharks (Carcharhinus melanopterus) swimming in shallow crystal clear water, Aldabra Atoll, Seychelles, Indian Ocean
American flamingo (Phoenicopterus ruber) feeding in water with hill in background. Floreana Island, Galapagos, Ecuador, June
Young Grey seal (Halichoerus grypus) exploring a seaweed garden in summer, Cairns Of Coll, Island of Coll, Inner Hebrides, Scotland, UK, North Atlantic Ocean, June
Alpine marmot (Marmota marmota), Hohe Tauern National Park, Austria, July
Roe Deer (Capreolus capreolus) doe running in a field of barley, Northumberland, England, UK, June. 2020VISION Book Plate
Black bear (Ursus americana) foraging for alpine berries during Autumn, on hillside, Cascade Mountains, Washington, USA
Guanaco (Lama guanicoe) herd below Torres del Paine, Torres del Paine National Park, Patagonia, Chile
Red grouse (Lagopus lagopus scoticus) on heather moorland showing habitat, Peak District National Park, UK, September
Red grouse (Lagopus lagopus scoticus) on heather moorland, Peak District NP, UK, September 2011. 2020VISION Exhibition. 2020VISION Book Plate
Two Arctic wolves (Canis lupus) in tundra landscape, Ellesmere Island, Nunavut, Canada, June 2008, Taken on location for BBC series, Frozen Planet, Summer
Red Fox (Vulpes vulpes) in distance in snow habitat. Vosges, France, December
Chamois (Rupicapra rupicapra) running through snowy forest. Vosges mountain, France, November
Dromedary Camel (Camelus dromedarius) on Erg Chebbi Dunes. Sahara Desert, Morocco, North Africa, March 2011
Mute swan (Cygnus olor) four on water in winter dawn mist, Loch Insh, Cairngorms NP, Highlands, Scotland UK, December. 2020VISION Exhibition. 2020VISION Book Plate
Long-tailed / Crab-eating macaque (Macaca fascicularis) foraging on coastline at low tide, Bako National Park, Sarawak, Borneo, Malaysia
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Long-tailed / Crab-eating macaque (Macaca fascicularis) foraging on coastline at dawn, Bako National Park, Sarawak, Borneo, Malaysia
Polar bear (Ursus maritimus) female and her two cubs on large iceberg off Baffin Island. Nunavut, Canada, August 2010
Female European moose (Alces alces) in flowering field, Elk, Morko, Sormland, Sweden, July 2009
Grey seal (Halichoerus grypus) on beach with wind blowing sand, Donna Nook, Lincolnshire, UK, November 2008
King Penguin (Aptenodytes patagonicus) male and female rear view looking out over colony, Saint Andrews Bay, South Georgia
Brown bear {Ursus arctos} near photographers wooden cabin, Valley of the Geysers, Kronotsky Zapovednik, Kamchatka, Far East Russia
Brown bear {Ursus arctos} at the Stained glass window, Valley of the Geysers, Kronotsky Zapovednik, Kamchatka, Far East Russia
Brown bear {Ursus arctos} at the Stained glass window in the Valley of the GeysersBrown bear {Ursus arctos} at the " Stained glass window" in the Valley of the Geysers, Kronotsky Zapovednik, Kamchatka, Far East Russia
Brown bear {Ursus arctos} at the Malachite Grotto, Valley of the Geysers, Kronotsky Zapovednik, Kamchatka, Far East Russia
Brown bear {Ursus arctos} fishing for salmon in the Kronotskaya River at the foot of the Kronotsky Volcano, covered with a fresh coat of snow in September, Kronotsky Zapovednik, Kamchatka
Stellers sea eagles {Haliaeetus pelagicus} gather beside Kuril Lake with Ilinsky volcano in the background, Kamchatka, Far East Russia, winter 2007
