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Elephants (Loxodonta africana) at waterhole drinking at night. One spraying water from trunk, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
African elephant (Loxodonta africana) charging through water Etosha National Park, Namibia
Long-tailed tit (Aegithalos caudatus caudatus) with feather for nest, Finland. April
Caribbean Flamingo (Phoenicopterus ruber) building up mud whilst brooding egg, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June
Landscape modified by the erosion process. Goreme National Park and the Rock Sites of Cappadocia UNESCO World Heritage Site. Turkey. December 2006
Pinnacles, also known as fairy chimneys or hoodoo, Love Valley. Goreme National Park and the Rock Sites of Cappadocia UNESCO World Heritage Site. Turkey. December 2006
Marine iguana (Amblyrhynchus cristatus) feeding underwater on algae. Endemic species. With Schooling black striped salema (Xenocys jessiae)(endemic species)
Willow (Salix) plantation, Cuenca, Castilla-La Mancha, Spain, December
African buffalo (Syncerus caffer) at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Hippopotamus (Hippopotamus amphibius) at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Fallow deer (Dama dama) at night, Gyulaj, Hungary Third place in the Nature Portfolio category of the World Press Photo Awards 2017
African buffalo (Syncerus caffer) and Warthog (Phacochoerus africanus) at night with full moon, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Great crested grebe (Podiceps cristatus) perfectly mimicking each others movements in courtship behaviour. Den Oever, The Netherlands May
Meerkats (Suricata suricatta), Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Great crested grebe (Podiceps cristatus) family on water, with adult feeding its young, Den Oever, The Netherlands, July
Early bumblebee (Bombus pratorum), visiting flax flowers, (Linum usitatissimum), Monmouthshire Wales, UK, May
Early bumblebee (Bombus pratorum) flying to feed on Purple toadflax (Linaria purpurea) Monmouthshire, Wales, UK. May
Peringueys desert adder (Bitis peringueyi) hidden in sand, Namibia
Laysan albatross (Phoebastria immutabilis), chick, Eastern island, Midway Atoll National Wildlife Refuge, Hawaii
Emperor penguin (Aptenodytes forsteri), chicks on ice, Snow Hill Island, Antarctic Peninsula
Puna / James flamingo (Phoenicoparrus jamesi) flock on Laguna Colorado, Reserva Eduardo Avaroa, altiplano, Bolivia September
Great Frigatebird (Fregata minor) chick in nest at sunset, San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
African buffalo (Syncerus caffer) with Yellow-billed oxpecker (Buphagus africanus) grooming, Chobe National Park Botswana
Hanuman Langurs (Semnopithecus entellus) in front of cenotaphs, Mandore Garden, Jodhpur, India. March 2015
Hanuman Langurs (Semnopithecus entellus) social grooming, Mandore Garden, Jodhpur, India
Hanuman Langurs (Semnopithecus entellus) group sitting in front of cenotaph, sunrise, Mandore Garden, Jodhpur, India. March 2015
Hanuman langurs (Semnopithecus entellus) grooming dog Mandore Garden, Jodhpur, India
African buffalo (Syncerus caffer) feeding on water lilies, Chobe River, Chobe National Park, Botswana. May
Marine iguana (Amblyrhynchus cristatus), male in full breeding colours Floreana Island, Galapagos
Honey bees (Apis mellifera) forming living bridge, Kiel, Germany, June
Deserta Grande wolf spider (Hogna ingens), Deserta Grande, Madeira, Portugal. Critically endangered
Giant panda (Ailuropoda melanoleuca) female, Huan Huan, holding baby age one month, Beauval Zoo, France. September 2017
Giant panda (Ailuropoda melanoleuca) female, Huan Huan, holding baby age three months, Beauval Zoo, France, November 2017
Whale headed / Shoebill (Balaeniceps rex) portrait. Swamps of Mabamba, Lake Victoria, Uganda
Short-finned pilot whale (Globicephala macrorhynchus) female carrying dead calf, with others swimming nearby, Los Gigantes, South Tenerife, Canary Islands, Atlantic Ocean, May
Short-finned pilot whale (Globicephala macrorhynchus) surfacing, South Tenerife, Canary Islands, Atlantic Ocean, July
Short-finned pilot whale (Globicephala macrorhynchus) female carrying dead calf, diving down from surface, Los Gigantes, South Tenerife, Canary Islands, Atlantic Ocean, May
Great hammerhead shark (Sphyrna mokarran) swimming over sandy seabed, South Bimini, Bahamas. The Bahamas National Shark Sanctuary, West Atlantic Ocean
Hoffmanns Two-toed sloth (Choloepus hoffmanni) mother and baby, aged 2 months, in tree, Costa Rica. Rescued and released by Aviarios Sloth Sanctuary
Sea otter (Enhydra lutris) mother and sleeping newborn pup (aged 3 days) Monterey, California, USA
Green sea turtle (Chelonia mydas) resting or sleeping on coral reef, Bonaire, Leeward Antilles, Caribbean region, Netherlands Antilles
Horned adder (Bitis caudalis) camouflaged in its environment, Namib Naukluft National Park, Namibia June
Asiatic Wild Buffalo (Bubalus arnee), swimming in lake at sunset. Kaziranga National Park UNESCO World Heritage Site, India. Feb 2016
Blackbuck (Antelope cervicapra), Male closeuphead portrait, Tal Chhapar Wildlife Sanctuary, Rajasthan, India
Humpback whale (Megaptera novaeangliae) breaching, Baja California, Mexico